Journal Pre-proof Micronuclei as biomarkers of DNA damage, aneuploidy, inducers of chromosomal hypermutation and as sources of pro-inﬂammatory DNA in humans Michael Fenech, Siegfried Knasmueller, Claudia Bolognesi, Nina Holland, Stefano Bonassi, Micheline Kirsch-Volders PII: S1383-5742(20)30062-4 DOI: https://doi.org/10.1016/j.mrrev.2020.108342 Reference: MUTREV 108342 To appear in: Mutation Research-Reviews in Mutation Research Received Date: 22 July 2020 Accepted Date: 19 October 2020 Please cite this article as: { doi: https://doi.org/ This is a PDF ﬁle of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the deﬁnitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its ﬁnal form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. © 2020 Published by Elsevier.