VOL. 10, NO. 7, JULY 2015 ISSN 1990-6145
ARPN Journal of Agricultural and Biological Science
© 2006-2015 Asian Research Publishing Network (ARPN). All rights reserved.
www.arpnjournals.com
247
EFFECT OF HUMIC ACID AND SULFUR ON GROWTH, SOME
BIOCHEMICAL CONSTITUENTS, YIELD AND YIELD
ATTRIBUTES OF FLAX GROWN UNDER NEWLY
RECLAIMED SANDY SOILS
Bakry A. B.
1
, Mervat Sh. Sadak
2
and El-karamany M. F.
1
Field Crops Research Department, National Research Centre, Dokki, Giza, Egypt
2
Department of Botany, National Research Centre, Dokki, Giza, Egypt
E-Mail: bakry_ahmed2004@yahoo.com
ABSTRACT
A field experiment was carried out at the experimental Station of National Research Centre, Al Nubaria district,
El-Behira Governorate-Egypt, in 2013/2014 and 2014/2015 winter seasons to investigate the influence of three sulfur
fertilizer levels and/or foliar humic acid on some morphological and biochemical parameters as well as yield quantity and
quality of flax plant grown under newly reclaimed sandy soil conditions. Results indicated that, humic acid foliar treatment
with 20 (mg/l) in addition to sulfur fertilizer at rate of (250 and 500 kg/fed) gave significant increases in seed, straw and oil
yield / fed. These increases due to the recorded increases in morphological criteria (plant height and root length, fresh and
dry weight of shoot and root), photosynthetic pigments, (chlorophyll a, chlorophyll b and carotenoids), carbohydrate
constituents (total carbohydrates, total soluble sugars and polysaccharides), IAA, phenol, free amino acids and proline
contents. Meanwhile, lipid peroxidation decreased significantly in response to the above mentioned treatment as compared
with control plants. Interaction between humic acid (20 cm/L) with sulfur at rate of (500 kg/fed) was the most effective
treatment as it gave the highest increases in all morphological criteria, biochemical parameters, yield and yield attributes
compared with the other treatments. Humic acid and sulfur fertilizer at all levels caused marked decreases in total saturated
fatty acids accompanied by marked increases in total unsaturated fatty acids. The essential fatty acids (Linoleic acid C 18:2
+ Linolenic acid C18:3) were increased by all applied treatments. It is worthy to mention that, the enhancement effects of
humic acid (20 mg/l) and sulfur fertilizer (500 kg/fed) were the most pronounced treatment on flax plant.
Keywords: fatty acid, flax, humic acid, lipid peroxidation, sulfur fertilizer, yields.
INTRODUCTION
Flax (Linum usitatissimum L.) is a source of two
products; flax seed for oil and fiber for linen products. In
Egypt, it is an old economic crop grown as a dual purpose
crop for seeds and fibers which is used for the
manufacture of linen. Linseed contains 30-48 % of oil
abundant in unsaturated fatty acids, especially α-linolenic
acid (40-68 %), where technical purity can reach 95%.
Linseed oil-chemical industry in drying oil (for the
production of printing and other inks, varnishes, paints and
linoleum).
Environmental pollution due to excessive
application of chemical fertilizer such as nitrogen fertilizer
to agricultural lands, surface waters and groundwater is
one of the most important environmental and social
concerns throughout the world especially in developing
countries (Parr et al., 1992). Soil organic matter has
beneficial effects on soil quality and positive effects on
crop productivity and also has the potential to sequester
carbon. In addition, organic matters could reduce the
application of industrial fertilizers in long term
(Rasmussen and Parton, 1994). Humic acid (HA) is one of
the main organic substances, which is an important
component of humic substances. Humic acid is complex
substances derived from organic matter decomposition.
Humic acid are the most significant constituents of organic
matter in both soils and municipal waste compost, and
have a relevant role in the cycling of many elements in the
environment and in soil ecological functions (Senesi et al.,
1996). According to previous investigations, humic seem
to have a particular favorable effect on the nutrient supply.
Foliar sprays of these substances also promote growth, and
increase yield and quality in a number of plant species
(Karakurt et al., 2009) at least partially through increasing
nutrient uptake, serving as a source of mineral plant
nutrients and regulator of their release (Atiyeh, et al.,
2002). Moreover, humic acid influence respiration-
process, the amount of sugars, amino acids and nitrate
accumulated (Boehme et al., 2005).
Sulfur (S) is an essential macroelement for plant
nutrition, but its concentration in plants is the lowest of all
the macronutrients. Sulfur is increasingly being recognized
as the fourth major essential nutrient element after
nitrogen (N), phosphorus (P), and potassium (K). Plants
are able to assimilate sulfate and reduce it to essential
amino acids, where S is involved in a range of metabolic
functions, including protein synthesis. Greater attention
needs to be paid to the role of S in balanced crop nutrition.
Sulfur plays an important role not only in the
growth and development of higher plants, but is also
associated with increased stress tolerance in plants (Nazar,
et al., 2011 and Osman and Rady, 2012). S deficiency
negatively affects the chlorophyll and N contents of leaves
and photosynthetic enzymes (Lunde, et al., 2008), and
consequently the reduction in yields and quality
parameters of crops (Hawkesford, 2000 and Osman and
Rady, 2012). Adequate S nutrition improves
photosynthesis and the growth of plants, and has