Effect of Divalent Metal Ions on Glucoamylase Activity of Glucoamylase
Isolated from Aspergillus niger
Okwuenu PC, Agbo KU, Ezugwu AL, Eze SO and Chilaka FC
Department of Biochemistry, Faculty of Biological Sciences, University of Nigeria, Nsukka, Enugu State, Nigeria.
Corresponding author: Okwuenu PC, Department of Biochemistry, Faculty of Biological Sciences, University of Nigeria, Nsukka, Enugu State, Nigeria, Tel:
+2348065251877; E-mail: prosperokwuenu@yahoo.com
Received date: Jan 06, 2017, Accepted date: Jan 28, 2017, Published date:Jan 31, 2017
Copyright: © 2017 Okwuenu PC, et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits
unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Abstract
This study investigates the production of glucoamylase from Aspergillus niger in a submerged fermentation
process using amylopectin fractionated from guinea corn starch as the carbon source. This work also studies the
effect of a few metal ions (Ca
2+
, Zn
2+
, Co
2+
, Fe
2+
, Mn
2+
and Pb
2+
) concentration on the glucoamylase activity. A
Fourteen day experimental study was carried out to determine the day of highest glucoamylase activity. Maximum
glucoamylase activity was observed on day five of the submerged fermentation; hence, day five was mass
produced. The specific activity for the crude enzyme was found to be 729.45 U/mg. The crude enzyme was purified
to the level of gel filtration (using sephadex G-100) via ammonium sulphate precipitation. Ammonium sulphate
saturation of 70% was found suitable to precipitate the enzyme. After ammonium sulphate precipitation and gel
filtration, the specific activities were found to be 65.98 U/mg and 180.52 U/mg respectively. The glucoamylase
activity was enhanced by 2 mM to 5 mM of Ca
2+
, Co
2+
, Fe
2+
, Mn
2+
and Zn
2+
but Pb
2+
had inhibitory effect on the
enzyme. The Michaelis constant, Km and maximum velocity Vmax of the enzyme was obtained from the
Lineweaver-Burk plot of initial velocity data at different substrate concentrations. They were found to be 770.75
mg/ml and 2500 µmol/min respectively, when using cassava starch as substrate. The enzyme glucoamylase is
known to have useful applications in food processing industries and fermentation biotechnology.
Keywords Glucoamylase; Enzyme; Aspergillus; Metal ions; Activity
Introduction
Glucoamylase (exo-1, 4-α-D-glucan-glucanohydrolase, EC 3.2.1.3)
belongs to the most important catalytically active proteins having
broad possibilities of technical use. It is an exo acting enzyme that
yields ß-D-glucose from the non-reducing chain ends of amylose and
amylopectin by hydrolyzing α-1, 4 linkages in a stepwise manner [1]. It
also hydrolyses α-1, 6 and the rare α-1, 3 linkages although at much
slower rate [2]. Generally, amylases, (that is α- amylases, β-amylases
and glucoamylase) can be produced either by submerged fermentation
(SmF) or solid state fermentation (SSF) procedures; however, the
convectional amylase production is carried out by submerged
fermentation [3] Glucoamylase production from microbial sources
especially from Aspergillus niger is generally extracellular, and the
enzyme can be recovered from culture fltrates. However, the extensive
utilization of glucoamylase is obtained by using the fungus Aspergillus
niger in enzyme production industries. Enzymes may require metal
ions for their maximal catalytic activity and are termed as
holoenzymes. Metals are responsible for right orientation of active site
of holoenzymes. Metal binding to enzyme is one of the factors
responsible for protein stabilization [4,5]. As a result of the important
role of glucoamylase, it is highly desirable to increase or enhance the
glucoamylase activity in other to improve its useful applications and
utilization.
Materials and methods
Chemicals
Bovine serum albumin (BSA), Sephadex G-100 was purchase from
Sigma Chemical Company Limited (USA). Folin–Ciocalteau was
purchased from Sigma-Aldrich (USA). Ammonium Sulphate and Tris
HCL salt were purchased from British Drug House (BDH) Chemicals
Limited (USA). All other chemicals used in this work were of
analytical grade and were obtained from reputable sources.
Collection of plant material and processing of the starch
Plant material (guinea corn seed and cassava tuber) was purchased
from the Ogige market in Nsukka, Enugu state, Nigeria, and its starch
was processed as described by Agboola [6].
Fractionation of starch into amylopectin
Te starch obtained from guinea corn was fractionated according to
the method of Sobukola and Aboderin [7].
Isolation and identifcation of the glucoamylase producing
fungi
Glucoamylase producing fungi was isolated and stored according to
the method described by Martin et al. [8] and the identifcation was
carried out using the method of Barnett and Hunter [9].
Okwuenu et al., Ferment Technol 2017, 6:1
DOI: 10.4172/2167-7972.1000141
Research Article Open Access
Ferment Technol, an open access journal
ISSN:2167-7972
Volume 6 • Issue 1 • 1000141
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ISSN: 2167-7972
Fermentation Technology