Molecular Ecology Resources (2008) 8, 919–922 doi: 10.1111/j.1755-0998.2008.02116.x
© 2008 The Authors
Journal compilation © 2008 Blackwell Publishing Ltd
Blackwell Publishing Ltd
PERMANENT GENETIC RESOURCES
Isolation and characterization of microsatellite loci from
the invasive ant Pheidole megacephala
DENIS FOURNIER, DAMIEN DUBOIS and SERGE ARON
Behavioural and Evolutionary Ecology CP 160/12, Université Libre de Bruxelles, Avenue F.D. Roosevelt, 50, B-1050 Brussels, Belgium
Abstract
We report the characterization of eight microsatellite markers in the big-headed ant Pheidole
megacephala, a pest ant registered in the list of ‘100 of the world’s worst invasive alien
species’. An enrichment protocol was used to isolate microsatellite loci, and polymorphism
was explored with 36 individuals collected in an invasive population from Australia and
20 individuals collected in a population from the native mainland location in South Africa.
These primers showed a number of alleles per locus ranging from two to 10, and expected
heterozygosities ranging from 0.083 to 0.826. Moreover, results of cross-species amplifica-
tion are reported in five other Pheidole species and in seven other ants of the subfamily
Myrmicinae.
Keywords: big-headed ant, colony structure, invasive species, microsatellite, Pheidole megacephala,
population structure
Received 24 October 2007; revision accepted 19 December 2007
Ants are popular organisms to address a variety of questions
in ethology, genetics, ecology, evolution or conservation
biology. Among the family Formicidae, Pheidole is one of
the most speciose and widespread genera: 898 described
species are distributed over the eight zoogeographical regions
where ant fauna are found (Neotropical, Nearctic, Palearctic,
Afrotropical, Malagasy, Oriental, Indo-Australian and
Australasian) (Hölldobler & Wilson 1990; Bolton 1995). In
the New World, species of the genus Pheidole are the most
abundant and diverse ants, and range from the northern
USA to Argentina (Wilson 2003). Pheidole species have
been studied for various aspects of their biology, including
the ecological correlates of their temporal and physical
castes and the associated division of labour (e.g. P. dentata,
Calabi & Traniello 1987; P. morrisi, Brown & Traniello 1998);
their highly biased colony sex ratio (P. desertorum, Helms 1999;
P. pallidula, Fournier et al. 2003); their foraging strategies
and territorial behaviour (e.g. P. pallidula, Passera et al.
1996; P. xerophylla (previously P. tucsonica) and P. gilvescens,
Langen et al. 2000; Tripet et al. 2006); and their impact on
biodiversity (e.g. P. megacephala, Passera 1994 and references
therein; Wetterer 2007; P. fervens and P. moerens, Garrison 1996).
The big-headed ant P. megacephala is well known as a
household and agricultural pest, and its negative ecological
impact on biodiversity may be greater than any other invasive
ant (Wetterer 2007). The species is nominated as one of
the 100 of the world’s worst invaders by the IUCN Species
Survival Commission Invasive Species Specialist Group
(Lowe et al. 2001). However, despite its ecological and
economical impacts, no genetic studies have been conducted
to decipher the evolutionary processes associated with
the invasion of the big-headed ant. Here, we describe the
development of eight microsatellite loci for this invasive
species and their application to individuals from Africa,
the supposed native range of the species (Wilson & Taylor
1967), and Australia.
Samples were collected in March 2005 from four Australian
populations and stored in ethanol at –80 °C. Total genomic
DNA was isolated using a standard phenol–chloroform
extraction protocol (Sambrook & Russell 2001). An enriched
library was made by ecogenics GmbH from size-selected
genomic DNA ligated into SAULA/SAULB-linker (Armour
et al. 1994) and enriched by magnetic bead selection with
biotin-labelled (CT)
13
, (GT)
13
(GTAT)
7
and (GATA)
7
oligo-
nucleotide repeats (Gautschi et al . 2000a, b). Of 374 recombinant
colonies screened, 163 gave a positive signal after hybridiza-
tion. Plasmids from 64 positive clones were sequenced and
primers were designed for 16 microsatellite inserts.
Correspondence: Denis Fournier, Fax: +32 (0)2 650.24.45; E-mail:
Denis.Fournier@ulb.ac.be