Aquaculture Nutrition. 2018;1–8. wileyonlinelibrary.com/journal/anu
|
1 © 2018 John Wiley & Sons Ltd
1 | INTRODUCTION
The intensification of production in shrimp culture demands that at-
tention be given to the reared organisms’ welfare, integrated water
management systems that avoid or minimize the pollution of water
bodies (i.e., release of nitrogenous compounds) and greater controls
over potential infections by pathogens (Xie & Yu, 2007; Moss, Moss,
Arce, Lightner, & Lotz, 2012; Xu & Pan, 2013). All of these actions are
particularly important for rearing the Pacific white shrimp Litopenaeus
vannamei, one of the most marketed aquatic organisms worldwide (Xu
& Pan, 2013).
Biofloc technology systems (BFTs) provide a sustainable solution
that improves water quality while removing toxic nitrogen (ammo-
nia, nitrite), and they also provide the highest growth performance
through the feed supplements and digestive stimulants from biofloc
organisms (Xu & Pan, 2013). BFTs are rich in natural microorganisms
Received: 18 January 2017
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Accepted: 16 December 2017
DOI: 10.1111/anu.12663
ORIGINAL ARTICLE
Antioxidant effects of nanoencapsulated lipoic acid in tissues
and on the immune condition in haemolymph of Pacific white
shrimp Litopenaeus vannamei (Boone, 1931)
Á.C. da Silva Martins
1,2
| J. Artigas Flores
1
| C. Porto
1
| L.A. Romano
1,2,3
|
W. Wasielesky Junior
1,2,3
| S.S. Caldas
1,4
| E.G. Primel
1,4
| I. Külkamp-Guerreiro
5
|
J.M. Monserrat
1,2,6,7
1
Universidade Federal do Rio Grande—FURG,
Rio Grande, Brazil
2
Programa de Pós-graduação em
Aquicultura, FURG, Rio Grande, Brazil
3
Instituto de Oceanografia (IO), FURG, Rio
Grande, Brazil
4
Escola de Química e Alimentos (EQA), FURG,
Rio Grande, Brazil
5
Faculdade de Farmácia, Universidade Federal
do Rio Grande do Sul, UFRGS, Porto Alegre,
Brazil
6
Instituto de Ciências Biológicas (ICB), FURG,
Rio Grande, Brazil
7
Laboratório de Bioquímica Funcional de
Organismos Aquáticos (BIFOA), Centro de
Biotecnologica e Doenças de Organismos
Aquáticos (CBD), IO, FURG, Rio Grande, Brazil
Correspondence
José Maria Monserrat, Instituto de Ciências
Biológicas (ICB), Universidade Federal do Rio
Grande—FURG, Rio Grande (RS), Brazil.
Email: monserrat_jm@furg.br
Funding information
Conselho Nacional de Desenvolvimento
Científico e Tecnológico, Grant/Award
Number: DT 311694/2016-0, PQ
307880/2013-3 and PQ 310993/2013-0;
MCTI/CNPq/FNDCT, Grant/Award Number:
408921/2013-7
Abstract
The antioxidant and oxidative damage responses and immune parameters in shrimp,
Litopenaeus vannamei, reared in clear water (CW) and in a biofloc system (BFT) were
evaluated. The animals were given feed supplemented with either nanocapsules con-
taining the antioxidant lipoic acid (NCLA) or empty nanocapsules (NC). Antioxidant
and oxidative stress analyses were evaluated by measuring glutathione S-transferase
activity (GST), reduced glutathione (GSH) concentration and lipid peroxidation (TBARS)
in the gills, hepatopancreas and muscle. The immune parameters were then deter-
mined by the differential hemocyte count in the haemolymph. The final weight of the
shrimp reared in both CW and BFT and treated with NCLA increased (p < .05). The
shrimps reared in BFT showed increased GST activity in the gills and hepatopancreas
after NCLA treatment (p < .05). Higher GSH levels were observed in the hepatopan-
creas with NCLA supplementation in CW and BFT (p < .05). Decreased levels of
TBARS were observed in the gills and muscles of shrimp supplemented with NCLA and
maintained in CW (p < .05). A decrease in the percentage of hyaline hemocytes and an
increase in granular hemocytes were observed in shrimp treated with NCLA in CW
and BFT (p < .05). The antioxidant treatment induced positive responses in terms of
the zootechnical, biochemical and immunological parameters.
KEYWORDS
antioxidant defences, antioxidants supplements, biofloc system, nanotechnology, oxidative
damage, shrimp