Iranian Journal of Animal Biosystematics (IJAB)
Vol.19, No.1, 33-45, 2023
DOI: 10.22067/ijab.2022.79924.1046
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Corresponding Author: rasoul.karamiani@gmail.com ©2023 FERDOWSI UNIVERSITY OF MASHHAD, IRAN
THIS WORK IS LICENSED UNDER A CREATIVE COMMONS ATTRIBUTION 4.0 INTERNATIONAL LICENSE.
IJAB
ISSN: 1735-434X (print); 2423-4222 (online)
Systematics and distribution of the Leopard
Geckos Eublepharis Gray, 1827 (Sauria,
Eublepharidae): A review
Rasoul Karamiani
Department of Biology, Faculty of Science, Razi University, 6714967346 Kermanshah, Iran
(Received: 5 December 2022; Accepted: 29 December 2022)
Abstract
The eyelid bearing geckos Eublepharidae including six widely disjunct genera and 45 species which
are distributed in tropical to temperate habitats of Eurasia, North-Central America and Africa. One of
the eublepharid genera is the fat-tailed geckos of the genus Eublepharis Gray, 1827. The specific
characters of Eublepharis involves flat basioccipital bone, deep axial pockets (armpit), eyelids well
developed and movable, pupil vertical, dorsal small scales juxtaposed larger tubercles. The fat-tailed
geckos as a vicariant group, encompass seven secretive, and nocturnal species: E. angramainyu
Anderson and Leviton, 1966, E. fuscus Börner, 1974, E. hardwickii Gray, 1827, E. macularius (Blyth,
1854), E. satpuraensis Mirza, Sanap, Raju, Gawai and Ghadekar, 2014, E. pictus Mirza and
Gnaneswar, 2022), and E. turcmenicus Darevsky, 1977 which are distributed from Turkey through
the Iranian Plateau to India as following: south-eastern Turkey, Syria, Iraq, Iran, Pakistan,
Afghanistan, Turkmenistan, and north-eastern and central India in a variety of habitats from dry karst
topography regions with gypsum deposits and clay-gravel soil to stony foothills. The data used for
the distribution maps were based on all available bibliographic records, and personal field
observations. The results showed that among the three species located in Iran, E. angramainyu has
the most widespread distribution and habitat diversity, and E. macularius has a very limited
distribution in the border of Iran and Afghanistan. The range of species distribution depends on
various factors including behavioral mechanisms of superior habitat selection, interactions with other
living organisms and physico-chemical factors.
Key words: Eublepharis, Eyelid geckos, Eublepharidae, Distribution, Asia.
INTRODUCTION
The eyelid geckos of the Family Eublepharidae encompass six widely disjunct genera and 45 species: the
cat gecko, Aeluroscalabotes (1 sp.) from Indonesia (Kalimantan), Malaysia (Sabah, Sarawak, west
Malaysia), Singapore, Thailand (Seufer et al., 2005), Coleonyx (8 sp.) from Central and western North
America, Eublepharis (7 sp.), and Goniurosaurus (25 sp.) from Asia, Hemitheconyx (2 sp.) from east and
west Africa, and Holodactylus (2 sp.) from east Africa (Vitt and Caldwell, 2013; Mirza et al., 2014;
Agarwal et al., 2022; Mirza and Gnaneswar, 2022). Boulenger (1885) based on movable eyelids, smooth
digit, procoelous vertebrae, a single parietal bone separated Eublepharidae from the Gekkota lizards. Then
Gadow 1901 reduced taxonomic level to subfamily and believed that they did not form a monophyletic
REVIEW ARTICLE Open access