Regulation of Male Sex Hormone Levels by Soy Isoflavones in Rats
Mi-Ae Yi, Hyun M. Son, Jung-Soon Lee, Chong-Suk Kwon,
Jin Kyu Lim, Young Keun Yeo, Young Sik Park, and Jong-Sang Kim
Abstract: Several studies have suggested that soybean in-
take is associated with a lower risk of prostate cancer. How-
ever, the mechanism of prostate cancer prevention by
soybeans remains unclear. Because prostate cancer is re-
ported to have an association with an increased level of
dihydrotestosterone (DHT) and soybean isoflavones are
known to inhibit 5a-reductase, which is involved in the con-
version of testosterone to DHT, the effects of soybean extract
and isoflavones on the plasma levels of male sex hormones
were investigated using male rats. In Experiment I, Sprague-
Dawley rats were fed diets with and without soy flour; in Ex-
periment II, rats were fed diets containing 2% soy methanol
extract or 0.2% semipurified isoflavones or a control diet.
The study showed a reduction of plasma DHT along with an
increase in total plasma androgen in rats fed soy flour or
semipurified isoflavones for 1 wk. These results suggest that
soy isoflavone intake may reduce plasma DHT level.
Introduction
Epidemiological studies show a lower incidence of hor-
mone-related cancer in Asians, including Koreans, Japanese,
and Chinese, than in their Western counterparts (1,2). Most
Asians eat a low-fat, high-fiber diet that is a rich source of
phytoestrogens, and there is an extremely low incidence of
prostate cancer. The soy-based foods included in this diet
may play a role in decreasing the prevalence of prostate can-
cer in Asian communities (3,4). The isoflavone levels are
much higher in the prostatic fluid from Hong Kong and Chi-
nese men who consume soybeans than in Portuguese and
British men (3). In animal models, most studies investigat-
ing the effects of soybeans show a reduced tumorigenesis
(5). For instance, a soy-containing diet was reported to re-
duce the severity of prostatitis in rats (6) and prevent the de-
velopment of dysplastic lesions of the prostate in neonatal
diethylstilbestrol-treated mice (7). Recently, soy isoflavones
have been shown to inhibit the growth of transplantable hu-
man prostate carcinoma and tumor angiogenesis in mice,
thereby suggesting multiple interacting mechanisms in pros-
tate cancer inhibition (8). In its early stages, prostate cancer
is hormone dependent, with castration causing tumor regres-
sion (9). Meanwhile, isoflavones of soy origin may play a
major role in the promotion and growth of prostate cancer by
modulating the metabolism of sex hormones. Genistein and
daidzein, the main isoflavones in soybeans, are known to in-
hibit 5a-reductase in vitro (9). This enzyme is involved in
the conversion of testosterone to 5a-dihydrotestosterone
(DHT), the main prostatic androgen responsible for the de-
velopment of prostate cancer (1,10). Therefore, there is a
strong possibility that soy isoflavones generate a significant
effect on the DHT level, inasmuch as they are known to be
strong inhibitors of 5a-reductase.
This communication reports that soy flour and isofla-
vones tended to lower the level of plasma DHT with a lim-
ited effect on the total androgen level in rats. In addition, a
significant amount of isoflavones was retained in the plasma
from rats fed an isoflavone-supplemented diet.
Materials and Methods
Animals and Diets
Male Sprague-Dawley rats (Korean Experimental Ani-
mal Center, Umsung, South Korea; 200–300 g body wt)
were housed individually in stainless steel cages and divided
into two and three experimental groups for two rounds of ex-
periments with 8 or 10 rats per group. The animals were
maintained at 22 ± 2°C on a 12:12-h light-dark cycle (lights
on at 0700) with free access to food and water. The diet com-
positions used in Experiment I are presented in Table 1. Au-
toclaved soy flour was blended into the powdered diet at
442.7 g/kg diet. In Experiment II, the rats in the control
group were fed ad libitum an AIN-76 diet containing 200 g
of casein, 3 g of DL-methionine, 150 g of corn starch, 500 g
of sucrose, 50 g of cellulose, 50 g of corn oil, 35 g of mineral
mixture, 10 g of vitamin mixture, and 2 g of choline
bitartrate per kilogram diet. The treatment groups were fed
an AIN-76 diet in which the cornstarch was partially re-
NUTRITION AND CANCER, 42(2), 206–210
Copyright © 2002, Lawrence Erlbaum Associates, Inc.
J.-S. Lee, J. K. Lim, Y. K. Yeo, Y. S. Park, and J.-S. Kim are affiliated with the Department of Animal Science and Biotechnology, Kyungpook National
University, Taegu 702-701, South Korea. M.-A. Yi is and H. M. Son was affiliated with the School of Food Science, Inje University, Kimhae 621-749, South
Korea. C.-S. Kwon is affiliated with the Department of Food Science and Nutrition, Andong National University, Andong 760-380, South Korea.