Acta zool, cracov. 35(3): 573-579, Krakow, 29 Jan. 1993 Fossil and Recent Anisopodomorpha (Diptera, Oligoneura): family Cramptonomyiidae. Vladimir A. BLAGODEROV, Ewa KRZEMINSKA and Wieslaw KRZEMINSKI Received: 25 Nov. 1992 Accepted to publication: 28 Dec. 1992 BLAGODEROV V. A., KRZEMINSKA E., KRZEMINSKI W. 1993. Fossil and Recent Aniso- podomorpha (Diptera, Oligoneura): family Cramptonomyiidae. Acta zool. cracov. 35(3): 573-579. Abstract. The key for distinguishing the Recent and fossil genera of the family Cramp- tonomyiidae is presented. Three new fossil species belonging to two new genera are descri bed: Tega penna gen. et sp. n. from the Upper Jurassic of Mongolia, T. karatavica sp. n. from the Upper Jurassic of Kazakhstan and Pivus sarus gen. et sp. n. from the Lower Cretaceous of Transbaikalia (South Siberia). Key words: Diptera, Anisopodomorpha, Axymyiomorpha, Cramptonomylidae, Pachy- neuridae, Tega, Pivus, fossil, Jurassic, Cretaceous, Mongolia, Siberia. Vladimir A. BLAGODEROV, Paleontological Institute, Academy of Sciences, Profsoyus- naya ul., 123, 117647 Moscow, Russia. Ewa KRZEMINSKA, Wieslaw KRZEMINSKI, Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, ul. Slawkowska 17,31-016 Krakow, Poland. INTRODUCTION The family Cramptonomyiidaewas established by HENNIG (1969) within the infraor- der Bibionomorpha. KRIvosHEINA & MAMAEV (1970) placed this family between the Pachyneuridae + Hesperididae and Anisopodidae. WOOD (1981) transferred it, at the subfamily level, to the family Pachyneuridae. However, in our opinion, Pachyneuridae and Cramptonomyiidae represent two separate lineages, which is apparent when compar- ing their wing venation (Figs 1-5). Pachyneuridae should be included into the infraorder Axymyiomorpha, whose representatives are characterizedby four radial veins terminating in the wing margin, although the vein R3 is very short (long R3+4 is present, formed by fusion of proximal sections of R3 and R4). In the higher Axymyiomorpha one medial vein is entirely reduced and d cell open (KRZEMINSKA et al. 1992). Cramptonomyiidae exhibit the features characteristic for the infraorder Anisopodomorpha: all medial veins present and d cell closed. Unique for the family is the course of R4; this vein originates from Rs