World Applied Sciences Journal 12 (6): 751-757, 2011
ISSN 1818-4952
© IDOSI Publications, 2011
Corresponding Author: Hafiz Muhammad Nasir Iqbal, Enzyme Biotechnology Laboratory,
Department of Chemistry and Biochemistry, University of Agriculture,
Faisalabad-38040, Pakistan, Tel: +92-307-6715243, E-mail: nasir_pk99@hotmail.com.
751
Purification and Kinetic Parameters Characterization of an Alkaline Protease
Produced from Bacillus subtilis through Submerged Fermentation Technique
Ishtiaq Ahmed, Muhammad Anjum Zia and Hafiz Muhammad Nasir Iqbal
Enzyme Biotechnology Laboratory, Department of Chemistry and Biochemistry,
University of Agriculture, Faisalabad-38040, Pakistan
Abstract: This paper reported the purification and kinetic parameters characterization of an alkaline protease
produced from Bacillus subtilis through submerged fermentation process using rice husk as growth supporting
substrate (by-product of rice industry) collected from Pearl Rice Mill Hafiz Abad, Faisalabad. Bacillus subtilis
was cultured in fermentation medium under some pre-optimized growth conditions. Maximum alkaline protease
activity of 216±4.32U/mL was obtained when fermentation medium of rice husk was inoculated with 4%
(4mL/100mL) inoculum size, 7% substrate concentration at pH 11 for 48 h fermentation time period with 2%
molasses as additional supplement material for fermentation medium. An alkaline protease was purified 1.49-fold
with specific activity of 74.66 U/mg in comparison to crude enzyme extract using ammonium sulfate
precipitation, dialysis and Sephadex-G-100 column chromatography. The enzyme was shown to have a relative
low molecular weight of 27kDa by sodium dodecyl sulphate poly-acrylamide gel electrophoresis (SDS-PAGE).
The enzyme displayed 10 and 45°C as an optimum pH and temperature respectively. Using casein as substrate,
the enzyme showed maximum activity (V ) of 148U/mL with its corresponding K value of 58µM. Among
max M
activators/inhibitors EDTA and Ca gave enhancing effect on purified alkaline protease where as SDS, Tween-
2+
81, Na and Hg caused enzyme inhibition and inactivation to variable extents. The specific activity and
+ 2+
substrate affinity of this alkaline protease from Bacillus subtilis is greater than those of other reported Bacillus
sp; therefore, it was concluded that it may be potentially useful for industrial purposes.
Key words: Alkaline protease Bacillus subtilis Purification SDS-PAGE Characterization
INTRODUCTION attention in attempts to exploit their physiological and
Proteases (EC 3.4.21-24) are enzymes that hydrolyze of the most important groups of industrial enzymes used
proteins and form a bulky cluster of enzymes which are in detergent, protein, brewing, meat, photographic,
ubiquitous in nature and most central category of leather, dairy industries, pharmaceutical and food industry
enzymes from an industrial point of view. An extensive [7, 8]. They used in pharmaceutical and food industry for
range of microorganisms has great potential to produce peptide synthesis in leather industry for dehairing and in
alkaline proteases under suitable growth conditions [1]. detergent industry as an additive of detergent formulation
Bacteria are the most dominant group of alkaline protease [2, 9] to substitute currently used toxic chemicals, is a
producers with the genus Bacillus being the most relatively new development and has conferred added
prominent and serve as an ideal source of these enzymes biotechnological importance [6].
[2, 3] because of their rapid growth and limited space Alkaline proteases have traditionally detained the
required for their cultivation [1]. An ease with which they predominant share of the industrial enzyme market
can be genetically manipulated to generate new enzymes accounting for about 60-65% of total global sale of
with altered properties that is desirable for their diverse enzymes [6, 10, 11]. About 35% of total microbial enzymes
applications [4]. used in detergent industry are derived from bacterial
The vast diversity of proteases, in contrast to their sources and most of them produced by Bacillus sp.
mode of action and specificity, has attracted worldwide [12, 13]. B. subtilis is a highly favorable bacterium for
biotechnological application [5, 6]. Proteases are the one