Reprod Dom Anim. 2019;00:1–6. wileyonlinelibrary.com/journal/rda
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1 © 2019 Blackwell Verlag GmbH
Received: 8 July 2019
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Revised: 17 September 2019
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Accepted: 26 September 2019
DOI: 10.1111/rda.13573
ORIGINAL ARTICLE
Induction of superovulation using inhibin antiserum and
competence of embryo development in wild large Japanese
field mice ( Apodemus speciosus )
Kanna Meguro
1
| Kazuki Komatsu
1
| Takuya Ohdaira
1
| Naomi Nakagata
2
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Akifumi Nakata
3
| Manabu Fukumoto
4
| Tomisato Miura
5,6
| Hideaki Yamashiro
1
1
Graduate School of Science and
Technology, Niigata University, Niigata,
Japan
2
Center for Animal Resources and
Development (CARD), Kumamoto
University, Kumamoto, Japan
3
Division of Life Science, Hokkaido
University of Science, Hokkaido, Japan
4
Department of Molecular Pathology, Tokyo
Medical University, Tokyo, Japan
5
Department of Bioscience and Laboratory
Medicine, Graduate School of Health
Sciences, Hirosaki University, Hirosaki,
Japan
6
Department of Radiation Biology, Institute
of Radiation Emergency Medicine, Hirosaki
University, Hirosaki, Japan
Correspondence
Hideaki Yamashiro, Laboratory of Animal
Reproduction, Graduate School of Science
and Technology, Niigata University, Niigata
959‐2181, Japan.
Email: hyamashiro@agr.niigata‐u.ac.jp
Funding information
Japan Atomic Energy Agency; Ministry of
Education; Japan Society for the Promotion
of Science
Abstract
Seasonally, bred wild mice provide a unique bioresource, with high genetic diversity
that differs from wild‐derived mice and laboratory mice. This study aimed to establish
an alternative superovulation method using wild large Japanese field mice ( Apodemus
speciosus) as the model species. Specifically, we investigated how the application of
inhibin antiserum and equine chorionic gonadotropin (IASe) during both the repro-
ductive and non‐reproductive seasons impact the ovulation rate and competence of
embryo development after in vitro fertilization (IVF) with fresh and cryopreserved
sperm. When the wild mice were superovulated by injecting eCG followed by human
chorionic gonadotropin (hCG), few oocytes were collected during the reproductive
and non‐reproductive seasons. In comparison, the number of ovulated oocytes was
dramatically enhanced by the administration of IASe, followed by isolation of ovu-
lated oocytes 24 hr after 30 IU hCG administration. The IVF oocytes that were in
vitro cultured (IVC) with medium containing serum further developed to the 2‐ and/
or 4‐cell stage using both fresh and frozen‐thawed sperm. In conclusion, we success-
fully established an alternative protocol for collecting ovulated oocytes from wild
large Japanese field mice by administering IASe and hCG during both the reproduc-
tive and non‐reproductive seasons. This study is the first to develop IVF–IVC wild
large Japanese field mice beyond the 2‐ and/or 4‐cell stage in vitro using fresh and
cryopreserved sperm. This approach could be used in other species of wild or endan-
gered mice to reduce the number of animals used for experiments, or in maintaining
stocks of germ cells or embryos.
KEYWORDS
embryo development, inhibin antiserum, sperm, superovulation, wild mice
1 | INTRODUCTION
The genetic diversity of laboratory strains of mice is limited in terms
of individual variation, with the original behavioural phenotypes of
wild mice no longer being expressed in laboratory mice (Koide et
al., 2011). Thus, using wild‐derived mice strains, including outbred
rodents, is advantageous for studying novel and diverse behavioural
phenotypes (Choi & He, 2015).
Recently, we reported that morphological and histological
changes to the testes of wild large Japanese field mice ( Apodemus
speciosus) are associated with seasonal changes (Akiyama et al.,
2015). Seasonally, bred wild mice provide a unique bioresource,