Defining reintroduction success using IUCN criteria for
threatened species: a demographic assessment
A. Robert
1
, B. Colas
2
, I. Guigon
1
, C. Kerbiriou
3
, J-B. Mihoub
4
, M. Saint-Jalme
1
& F. Sarrazin
3
1 CESCO, UMR 7204, MNHN-CNRS-UPMC, Muséum National d’Histoire Naturelle, Paris, France
2 Univ. Paris Diderot, Sorbonne Paris Cité, ESE, UMR 8079, UPS-CNRS-AgroParisTech, Univ. Paris Sud 11, Orsay, France
3 CESCO, UMR 7204, MNHN-CNRS-UPMC, Université Pierre et Marie Curie, Paris, France
4 UFZ – Helmholtz Centre for Environmental Research, Leipzig, Germany
Keywords
reintroductions; conservation translocations;
population dynamics; extinction; IUCN;
population viability analysis; threatened
species.
Correspondence
Alexandre Robert, CESCO, UMR 7204,
MNHN-CNRS-UPMC, Muséum National
d’Histoire Naturelle, 55, Rue Buffon, 75005
Paris, France. Tel: 33 1 40 79 57 27; Fax:
33 1 40 79 38 35; Email: arobert@mnhn.fr
Editor: Iain Gordon
Associate Editor: Rob Slotow
Received 17 July 2014; accepted 25
November 2014
doi:10.1111/acv.12188
Abstract
Despite recent efforts to develop the science of reintroduction biology, there is still
no general and broadly accepted definition of reintroduction success. We investi-
gate this issue based on the postulates (1) that successful reintroduction programs
should produce viable populations and (2) that reliable assessments of ultimate
success require that populations have reached their regulation phase. We assessed
if the viability of these reintroduced populations could be evaluated using the same
criteria as for remnant populations, such as the Internation Union for Conserva-
tion of Nature (IUCN) Red List criteria. Using modeling, we projected the viabili-
ties of theoretical populations with various life history and environmental
characteristics and we tested whether population sizes (criterion D of the IUCN)
and other potential predictors are relevant proxies of the risk of extinction (crite-
rion E of the IUCN) in the case of remnant populations with an unknown past
history and in the case of reintroduced populations that have reached their car-
rying capacity. We found that, as for remnant populations, population size can be
used as a relevant indicator (although subject to considerable uncertainty) of the
viability of reintroduced populations. However, the results demonstrate the
importance of the reintroduction failure filter, that is, the fact that the reintro-
duced populations that have successfully reached their carrying capacity are those
with the highest and more stable growth rates, especially if populations have been
reintroduced with a few individuals. As a consequence, the general relationship
between the current size of a population and its projected viability will, most
likely, differ considerably between remnant and reintroduced populations.
Overall, our results demonstrate that there are no theoretical limitations on the
application of some of the criteria widely used for remnant populations to define
reintroduction success, although these criteria are very conservative for reintro-
duced populations and might be rescaled to account for the demographic filter
that early extinction constitutes for these populations.
Introduction
Reintroduction research has long been based on descriptive
or opportunistic studies (Seddon, Armstrong & Maloney,
2007), but attempts are underway to adopt a more integra-
tive, standardized and theoretically sound approach to
reintroductions and other conservation translocations
(e.g. assisted colonizations; IUCN/SSC, 2013) to improve
project outcomes and contribute to fundamental ecological
research (Sarrazin & Barbault, 1996; Armstrong & Seddon,
2008). One particularly important issue is the quantification
of the roles of the various intrinsic, environmental and man-
agement factors on conservation translocation success,
which in turn requires a standardized definition of success.
However, there are still no general and broadly accepted
success criteria (Sarrazin & Barbault, 1996; Fischer &
Lindenmayer, 2000; Sutherland et al., 2010). Most authors
consider that reintroduction programs should aim at estab-
lishing viable populations (Armstrong & Seddon, 2008;
Schaub et al., 2009; IUCN/SSC, 2013). However, while rein-
troduction failures are generally easy to identify (e.g. docu-
mented extinction of the released stock and their offspring),
neither conceptual framework nor criteria are unanimously
used to define the bounds of viability.
The notion of population viability is, nevertheless, at the
core of conservation research and long-term viability is one
of the most universal and important concepts considered to
define the conservation status of remnant populations
(Beissinger & McCullough, 2002). Several conservation
agencies have developed assessment protocols based on
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