Int. J. Pharm. Sci. Rev. Res., 30(1), January – February 2015; Article No. 60, Pages: 340-344 ISSN 0976 – 044X
International Journal of Pharmaceutical Sciences Review and Research
Available online at www.globalresearchonline.net
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340
Saleha Y. M. Alakilli
Department of Biology, King Abdulaziz University, Jeddah/ Northern Border University, Arar, Saudi Arabia.
*Corresponding author’s E-mail: salakilli@yahoo.co.uk
Accepted on: 20-11-2014; Finalized on: 31-12-2014.
ABSTRACT
With the recent advances in molecular biology techniques, a great interest focused towards the improving livestock production in
general and meat production in specific. In the present study is to identify the genetic polymorphism of calpastatin and callipyge
genes (CAST and Clpy) related to meat production in two Saudi sheep breeds (Najdi and Harri). The detection was done using the
restriction fragment length polymorphism for the polymerase chain reaction products (PCR-RFLPs). Genomic DNA was isolated from
20 animals from each breed; The PCR products were digested with MspI restriction enzyme for calpastatin gene and FaqI restriction
enzyme for callipyge gene. The results indicated the presence of polymorphism in the callipyge gene was detected homozygous
genotype AA – 1.00 only in the two breeds understudy. The heterozygous genotype AG and homozygous genotype GG was not
detected. Calpastatin locus digested with MspI had two genotypes MM and MN. The highest allelic frequency was for allele M allele
but homozygous genotype NN has not been observed. The CAST locus had two genotypes MM (0.64 and 0.26 from Najdi and Harri)
and MN (0.36 and 0.74 from Najdi and Harri), but homozygous genotype NN has not been observed. It may be concluded that CLPG
locus is a monomorphic for population sheep kept in Saudi. The locus of CAST gene is polymorphic for breeds of sheep involved
Najdi and Harri. Najdi sheep breed showed the highest observed heterozygosity for CAST gene.
Keywords: Sheep, PCR-RFLP, Calpastatin gene, Callipyge gene.
INTRODUCTION
t the present time, the consumers demand for
sheep meat is not focused on quantitative traits
only, but also qualitative traits of meat too. Animal
breeders could accelerate the rate of genetic
improvement attained in carcass composition and meat
quality traits through the application of gene-assisted
selection (GAS), based simply on incorporating some
candidate genes in traditional breeding programs
1
.
The Najdi or Nejdi is a breed of domestic sheep native to
the Najd region of the Arabian Peninsula. Though it is
primarily raised in Saudi Arabia, Nadji sheep are also
present in Kuwait, Jordan, Oman, and Iraq
2
. The Najdi has
a distinctive appearance that has even been celebrated in
Saudi “sheep beauty pageants” not unlike livestock shows
and sales in the West. They are a very tall breed,
averaging 76-86 centimeters (30-34 inches) in height at
the withers
2
. They have long, Roman nosed faces with
drooping ears. Ewes are polled and rams may be either
polled or have scars.
They are generally black with white faces and white on
the legs and tail. Top Najdi ewes can sell for 20,000-
30,000 Saudi riyals ($5,300-$8,000 USD), while rams
which can sire many more offspring can fetch hundreds of
thousands
2
.
Najdi are highly adapted to life in desert conditions,
though it is less drought tolerant than some breeds, such
as the Awassi
3,4
. Though its meat may be consumed
locally, it is especially valued for its milk and long, straight
wool.
There are many of published articles on two genes
associated with meat-related traits in varied sheep
breeds, CAST and Callipyge genes
5
.
Calpastatin (CAST) gene is located on the fifth
chromosome of sheep and plays important roles in
formation of muscles, degradation and meat tenderness
after slaughtering
6
. It was also proved that there is a
relationship between polymorphism in the calpastatin
gene in sheep and slaughter traits such as lamb’s body
weight at birth and its growth rate until weaning
7
.
Associations between variation in CAST and carcass and
meat quality traits in sheep, there was also a genetic
variation in the CAST gene
8-11
The Callipyge (CLPG) gene is most documented gene for
double muscle in sheep which causes a postnatal muscle
hypertrophy that is localized to the pelvic limbs and
loin
12
. The mutation cognizant of unique muscling
phenotype in sheep was first discovered in 1983 in
Oklahoma in Dorset breed. The locus of CLPG gene was
mapped to the telomeric region of ovine chromosome
18
13
. The CLPG gene has unique inheritance pattern,
which has been termed as polar overdominance
14
. The
term polar overdominance was used to describe the
genetic model in which only heterozygous individuals that
were receiving the newly identified allele from their sire
expressed the unique phenotype
14
. Callipygephenotypic
effects were previously estimated in heterozygous lamb
but the mutant CLPG allele have to be inherited from sire
and normal allele have to be inherited from dam. The
others combination of allele can be classified as non
carrier lambs with normal phenotype
15
.
Analysis of Polymorphism of Caplstatin and Callipyge Genes in Saudi Sheep Breeds
Using PCR-RFLP Technique
A
Research Article