Abstract Basal keratins, suprabasal keratins, filag-
grin, and cornified cell envelope (CCE) precursor pro-
teins are expressed during the differentiation of epi-
dermal keratinocytes. These molecules are coordi-
nately expressed during epidermal differentiation. The
present study investigated the expression patterns of
keratins and CCE precursor proteins in 15 patients
with epidermolysis bullosa simplex (EBS), which is
caused by mutations in the genes that encode for the
basal keratins, keratins 5 and 14. The patterns of ex-
pression of keratins 5, 14, 1 and 10, filaggrin, and of the
three major CCE precursor proteins, involucrin, lori-
crin and small proline-rich proteins 1 and 2 (SPRs),
were studied immunohistochemically and by electron
microscopy. In 14 of the 15 patients with EBS, the dis-
tribution pattern of keratins was not altered. In one
neonate with EBS, basal cell keratins were expressed
in the suprabasal layers. Ultrastructurally, numerous
clumped tonofilaments were observed in the basal and
suprabasal cells. In all cases, findings were positive for
filaggrin in the granular cells, with positivity for in-
volucrin in the upper spinous and granular cells. The
upper spinous cells and granular cells were positive
for SPRs 1 and 2, and loricrin was expressed in granu-
lar cells. Ultrastructurally, no marked abnormality was
observed in the suprabasal layers such as a decrease
in, or agglutination of, keratin filaments, except in one
neonate. A CCE about 15 nm thick was formed nor-
mally in the cell membrane of cornified cells. The pat-
terns of distributions of basal cell keratins, suprabasal
keratins, filaggrin, and CCE precursor proteins, as well
as the ultrastructural findings, resembled those of nor-
mal skin. Thus, the abnormality in basal cell keratins
in patients with EBS did not appear to alter the pat-
terns of expression of the keratins and CCE precursor
proteins.
Key words Keratin · Epidermolysis bullosa simplex ·
Cornified cell envelope
Introduction
The keratins, which are the major structural proteins of
the keratinocytes, are important in maintaining the func-
tion of the skin. The process of keratinization is initiated
by the expression of keratins 5 (K5) and 14 (K14) in the
basal cells, and terminates with the production of horny
cells in which filaggrin and the cornified cell envelope
(CCE) precursor proteins play important roles. Thus, it
seems that mutations in the basal keratin gene could lead
to the abnormal expression of keratin and the filaggrin fil-
ament network that could lead to a defect in CCE produc-
tion. Based on electron microscopic finding of an abnor-
mal organization of keratin intermediate filaments in cul-
tured epidermal keratinocytes of epidermolysis bullosa
simplex (EBS) [18], mutations of the keratin gene in pa-
tients with EBS were first reported in 1991 [1, 7]. Many
other groups have since reported similar gene mutations
[2, 3, 5, 6, 10, 11, 19, 21, 29]. However, only a few stud-
Yuko Sasaki · Hiroshi Shimizu · Masashi Akiyama ·
Kozo Yoneda · Akemi Ishida-Yamamoto ·
Shinichi Watanabe · Junichi Hata · Takeji Nishikawa
Abnormalities of basal cell keratin in epidermolysis bullosa simplex
do not affect the expression patterns of suprabasal keratins
and cornified cell envelope proteins
Arch Dermatol Res (1998) 290 : 591–597 © Springer-Verlag 1998
Received: 8 April 1998 / Received after revision: 17 July 1998 / Accepted: 30 July 1998
ORIGINAL PAPER
Y. Sasaki (Y) · H. Shimizu · T. Nishikawa
Department of Dermatology,
Keio University School of Medicine, 35 Shinanomachi,
Shinjuku, Tokyo 160-8582, Japan
Tel. +81-3-3353-1211, ext. 2413; Fax +81-3-3351-6880;
e-mail: yuko@mc.med.keio.ac.jp
M. Akiyama
Division of Dermatology, Kitasato Institute Hospital,
Tokyo, Japan
K. Yoneda
Department of Dermatology,
Kyoto University Faculty of Medicine, Kyoto, Japan
A. Ishida-Yamamoto
Department of Dermatology, Asahikawa Medical College,
Asahikawa, Japan
S. Watanabe
Department of Dermatology,
Teikyo University School of Medicine, Tokyo, Japan
J. Hata
Department of Pathology, Keio University School of Medicine,
Tokyo, Japan