Journal of Phytopathology. 2017;1–7. wileyonlinelibrary.com/journal/jph
|
1 © 2017 Blackwell Verlag GmbH
Received: 15 May 2017
|
Accepted: 26 July 2017
DOI: 10.1111/jph.12615
ORIGINAL ARTICLE
Phenylalanine ammonia-lyase gene induction with
benzothiadiazole elevates defence against Lasiodiplodia
theobromae in tea in India
Shibu Das
1
| Prosenjit Chakraborty
1
| Parimal Mandal
2
| Dipanwita Saha
3
|
Aniruddha Saha
1
1
Department of Botany, University of North
Bengal, Siliguri, India
2
Department of Botany, Raiganj University,
Raiganj, India
3
Department of Biotechnology, University of
North Bengal, Siliguri, India
Correspondence
Aniruddha Saha, Department of Botany,
University of North Bengal, Siliguri, India.
Email: asahanbu@yahoo.co.in
Funding information
UGC
Abstract
Tea is widely used as an aromatic beverage cultivated in more than 35 countries from
different continents. The global production of tea during the year 2015–2016 was
5,303 million kg where India stands 2nd contributing 23% of the world production.
Diplodia disease is one of the most common diseases of tea in sub-Himalayan West
Bengal, India. Phenylpropanoids act as a precursor of several secondary metabolites
that are synthesized at the time of biotic and abiotic stress conditions. During the in-
duction of systemic acquired resistance, salicylic acid production is regulated by phe-
nylalanine ammonia-lyase (PAL) gene expression. In this communication,
twelve-month-old tea seedlings (TV-26) were treated with benzothiadiazole (BTH) and
inoculated with Lasiodiplodia theobromae. Transcript accumulation in different treat-
ments, that is treated-uninoculated, untreated-inoculated and treated-inoculated were
measured in comparison with untreated-uninoculated control to study the effect of
BTH in defence induction. From the results, it was evident that BTH-treated and
L. theobromae-inoculated plants showed higher transcript accumulation (3.81 μg/μl)
and reduced disease index (3.8 ± 0.02) in comparison with untreated control where
transcript accumulation was 3.26 μg/μl and disease index reduced to 14.6 ± 0.05. After
sequencing, nucleotide and deduced protein sequences were compared, and 3D struc-
ture was established where three lyase superfamily (cl26059) motifs were found. The
increase in PAL activity was also observed in treated-inoculated plants (from 2.5 to
4.4 μmol min
−1
g
−1
) in comparison with healthy control (from 2.5 to 2.7 μmol min
−1
g
−1
)
after 4th day of treatment. Nonetheless, BTH might be one of the significant inducers
of PAL which can be used to induce of disease resistance in tea plants in India.
KEYWORDS
benzothiadiazole, diplodia disease, phenylpropanoids, transcript accumulation
1 | INTRODUCTION
Tea (Camellia sinensis (L.) O. Kuntze.) is a tree member of the fam-
ily Theaceae. Processed tea leaves are widely used as an aromatic
beverage cultivated in more than 35 countries from different conti-
nents. It is a rich source of antioxidant, epigallocatechin, epicatechin,
gallocatechin and catechin and several other phenolic compounds that
play important roles in improving human health and also providing re-
sistance against a wide range of pathogens. Green leaves of tea are also
used as vegetable in Burma, Thailand and also have some medicinal
importance against blood pressure, coronary heart diseases and diabe-
tes (Azzahra, Fouzia, Mohammed, & Noureddine, 2012; Chen, 1999;