Middle-East Journal of Scientific Research 3 (3): 134-139, 2008
ISSN 1990-9233
© IDOSI Publications, 2008
Corresponding Author: Ghaleb Adwan, Department of Biology and Biotechnology,
An-Najah N. University, P.O. Box 7, Nablus, Palestine
134
Synergistic Effects of Plant Extracts and Antibiotics on
Staphylococcus aureus Strains Isolated from Clinical Specimens
Ghaleb Adwan and Mohammad Mhanna
1 2
Department of Biology and Biotechnology, An-Najah N. University,
1
P.O. Box 7, Nablus, Palestine
Faculty of Graduate Studies, Department of Environmental Sciences,
2
An-Najah N. University, P.O. Box 7, Nablus, Palestine
Abstract: This study has been done to evaluate the interaction between water extracts of Psidium guajava,
Rosmarinus officinalis, Salvia fruticosa, Majorana syriaca, Ocimum basilucum, Syzygium aromaticum,
Laurus nobilis and Rosa damascena alone and then synergy testing of these extracts with known antimicrobial
agents of different mechanisms (protein synthesis inhibition: oxytetracycline HCl and gentamicin sulfate; cell
wall synthesis inhibition: penicillin G and cephalexin; folic acid synthesis inhibition: Sulfadimethoxine as
sodium; and nucleic acid synthesis inhibition: enrofloxacin) using both well-diffusion and microdilution method.
This study was conducted against five S. aureus isolates; one is Methicillin-resistant Staphylococcus aureus
(MRSA) and 4 Methicillin-sensitive Staphylococcus aureus (MSSA). The results of the conducted experiments
using well-diffusion method demonstrate that these plants showed in vitro interactions between antimicrobial
agents and plant extracts were additive against the five strains of S. aureus, while using microdilution method
showed synergistic effects between combination of antibiotics and plant extracts with significant reduction in
the MICs of the test antibiotics against these strains of S. aureus. This change in MIC was noticed in all plant
extracts against test antibiotics including these plants showed weak antibacterial activity by well diffusion
method. Also our results showed that synergism effect between antimicrobial agent and plant extract was
occurred in both sensitive and resistant strains but the magnitude of minimum fold inhibition in resistant strains
especially MRSA strain was higher than the sensitive strains.
Key words: Plant extracts Synergistic effects Antimicrobial agents Microdilution method Well
diffusion method
INTRODUCTION Few studies have found that the efficacy of
The wide use of antibiotics in the treatment of with crude plant extracts against different pathogens
bacterial infections has led to the emergence and spread including S. aureus, P. aeruginosa, E. coli, Extended
of resistant strains. Methicillin-resistant Staphylococcus Spectrum -lactamases-producing multidrug-resistant
aureus (MRSA) is a major cause of nosocomial infections. E. coli and vancomycin-resistant enterococci
MRSA infections are very difficult to cure because MRSA (Enterococus fecalis) [6-18]. Some Palestinian plants
strains are resistance against almost all clinically available exhibit significant potency against human bacterial
antibiotics. For most MRSA strains, glycopeptide-type [19-21]. But plant extracts as antimicrobials are rarely used
drugs such as vancomycin are the only effective as systemic antibiotics at present, this may be due to their
antimicrobial agents [1]. However, vancomycin-resistant low level of activity, especially against Gram-negative
S. aureus (VRSA) has been reported [2-4]. Thus, it is bacteria. Here we are trying to investigate an alternative
extremely important to find new antimicrobial agents or approach to the treatment of bacterial infections by able
new ways that are effective for the treatment of infectious to change the phenotype of a resistant pathogen to
diseases caused by drug-resistant bacteria including certain antibiotics to more susceptible pathogen to that
MRSA [5]. antibiotics. This alternative approach has been evaluated
antimicrobial agents can be improved by combining them