Vol.:(0123456789)
Environmental Science and Pollution Research (2024) 31:15571–15579
https://doi.org/10.1007/s11356-024-32117-3
RESEARCH ARTICLE
Diversity and genetic structure of freshwater shark Wallago attu:
an emerging species of commercial interest
Shahid Sherzada
1,2
· Nimra Hussain
2
· Ali Hussain
3
· Mohamed A. M. El‑Tabakh
4
· Saeed Akram Khan
1
Received: 18 April 2023 / Accepted: 17 January 2024 / Published online: 1 February 2024
© The Author(s), under exclusive licence to Springer-Verlag GmbH Germany, part of Springer Nature 2024
Abstract
Pakistan has natural freshwater resources acting as a hotspot for diverse fish fauna. However, this aquatic fauna is declining
at an alarming rate due to over-exploitation, habitat degradation, water pollution, climate change, and certain anthropogenic
activities. The freshwater shark, Wallago attu, is a popular edible catfish inhabiting these freshwater ecosystems. Habitat
degradation, overfishing, and human activities are heavily impacting the natural population of this species. So, sound knowl-
edge about its population structure is necessary for its proper management in natural waters. The current study involves
utilizing two mtDNA markers (COI, Cytb) to assess the genetic structure and differentiation among W. attu populations of
Pakistani Rivers. Genetic variability analysis indicated a high haplotype (0.343 ± 0.046–0.870 ± 0.023) and low nucleotide
diversity (0.0024 ± 0.012–0.0038 ± 0.018) among single and combined gene sequences, respectively. Overall, River Indus
was populated with more diverse fauna of Wallago attu as compared to River Chenab and River Ravi. Population pairwise,
F
st
values (0.40–0.61) were found to be significantly different (p < 0.01) among three Riverine populations based upon com-
bined gene sequences. The gene flow for the combined gene (COI + Cytb) dataset among three populations was less than 1.0.
The transition/transversion bias value R (0.58) was calculated for testing of neutral evolution, and it declared low genetic
polymorphism among natural riverine populations of Wallago attu. The current study’s findings would be meaningful in
planning the management and conservation of this economically important catfish in future.
Keywords Wallago attu · Genetic diversity · Mitochondrial DNA · COI · Cytb · Mitochondrial DNA; Genetic diversity;
Population structure
Introduction
Freshwater ecosystems are home to diverse aquatic flora
and fauna. These ecosystems are now facing threats due to
gradual loss of biodiversity. The physical, chemical, and
biological components of freshwater ecosystems are heavily
impacted by anthropogenic pressure (Navarro-Ortega et al.
2015). World Wide Fund for Nature (WWF) has reported
that the current rate of population decline in freshwater
fishes is two times more than marine water fish fauna (WWF
2016). For surveillance and conservation, genetic charac-
terization of natural fish populations is mandatory. Genetic-
based approaches are very valuable to better understand the
genomic basis of adaptive divergence in fish populations
(Kumari et al. 2017).
Wallago attu, commonly known as helicopter fish, fresh-
water shark, or Asian sheat catfish, is found in the natural
waters of South and Southeast Asia from Pakistan to Indo-
nesia. It is placed among the 10 largest catfish species in the
world (Hogan 2011). In Pakistan, it is locally named “Mul-
lee” or “Jarko” (Mirza 2003). Wallago attu inhabits rivers,
lakes, and reservoirs and is usually captured by traditional
fishing methods (Montana et al. 2011; Renjithkumar et al.
2016).
Responsible Editor: Bruno Nunes
* Shahid Sherzada
shahid.sherzada@gcu.edu.pk
1
Department of Zoology, Government College University
Lahore, Lahore, Pakistan
2
Department of Fisheries and Aquaculture, University
of Veterinary and Animal Sciences, Lahore, Pakistan
3
Institute of Zoology, University of the Punjab, Lahore,
Pakistan
4
Zoology Department, Faculty of Science, Al-Azhar
University, Cairo, Egypt