Introduction
Body colour plays a significant role in the lives of
amphibians, whose interaction with solar radiation has
important implications for their thermal biology and
correlated survival rate (e.g., Terentiev, 1950; Childs,
1953; Vences et al., 2002; Sanabria et al., 2014; Stuart-
Fox et al., 2017; Martínez-Freiría et al., 2020; Smith
et al., 2021). Thus, studying amphibian body colour
is useful to gain a more in-depth understanding of
the phenotypic plasticity and ecological adaptability
of these organisms (e.g., Boero, 2013; Kolenda et al.,
2017). In recent decades morphological abnormalities,
including those involving body colour, have become
a common issue for almost all amphibian populations
in Ukraine, apparently as a consequence of habitat
degradation (Henle et al., 2017a; Reshetylo et al., 2019;
Palamarenko, 2020), and it is important to document
these and recognise their origins.
Amphibians are known to be affected directly and
indirectly by exposure to pollutants or land-use changes,
which can result in internal and external morphological
abnormalities (Marushchak et al., 2017; Henle et al.,
2017a; Marushchak and Muravynets, 2018; Hegde et
al., 2019). Of these, colour anomalies are just one group
of skin morphology aberrations that were classified as
Type S1 (“skin malformations”) by Nekrasova (2008).
However, determining the origin of a particular anomaly
Herpetology Notes, volume 14: 1239-1251 (2021) (published online on 29 September 2021)
A GIS approach to the study of colour anomalies in amphibians
of Ukraine reveals the deleterious effect of human impacts
Oleksii Y. Marushchak
1,*
, Oksana D. Nekrasova
1,4
, Volodymyr M. Tytar
1
, Nazar A. Smirnov
2
, Oleksiy V.
Korshunov
3
, Mihails Pupins
4
, Galyna I. Mykytynets
5
, Arturs Skute
4
, Klaus Henle
6
, and Hinrich Kaiser
7
1
I.I. [Ivan Ivanovitch] Schmalhausen Institute of Zoology,
National Academy of Sciences of Ukraine, Bohdana
Khmelnytskogo Street 15, Kyiv 01030, Ukraine.
2
Chernivtsi Regional Local Lore Museum, Olga Kobylyans’koi
Street 28, Chernivtsi 58002, Ukraine.
3
Department of Zoology and Animal Ecology, V.N. [Vasily
Nazarovich] Karazin Kharkiv National University, Svobody
Square 4, Kharkiv 61022, Ukraine.
4
Department of Ecology, Institute of Life Sciences and
Technologies, Daugavpils Universitāte, Parādes iela 1a,
Daugavpils 5401, Latvia.
5
Pryazovsky National Nature Park, Ivan Bohun Street 46,
Melitopol 72300, Ukraine.
6
UFZ – Helmholtz Centre for Environmental Research,
Department of Conservation Biology & Social-Ecological
Systems, Permoserstraße 15, Leipzig 04318, Germany.
7
Department of Vertebrate Zoology, Zoologisches
Forschungsmuseum Alexander Koenig, Adenauerallee
160, 53113 Bonn, Germany; and Department of Biology,
Victor Valley College, 18422 Bear Valley Road, Victorville,
California 92395, USA.
*
Corresponding author. E-mail: ecopelobates@gmail.com
© 2021 by Herpetology Notes. Open Access by CC BY-NC-ND 4.0.
Abstract. Our study provides a review of colour anomalies in amphibians from Ukraine during the 20
th
and early 21
st
centuries.
Observations including melanism, flavinism, leucism, and blue axanthism were assembled from the published literature (1909–
2018) and during field surveys (2000–2017). Blue colouration was the most common abnormal variant (81.5%; n = 106), and
colour anomalies were recorded in 13 of Ukraine’s 24 administrative regions (oblasts), mainly along the Dnieper River and in
the Carpathian Mountains. The largest number of anomalies was found in the Poltava (26.5%) and Kyiv (20.4%) Oblasts. We
also explored the relationship between abnormal colouration and environmental variables using a GIS framework. Correlations
existed mainly with geographic location and temperature-related parameters (e.g., reference evapotranspiration), as could
be expected for interactions of morphological or physiological anomalies. However, the Human Footprint, an integrated
index of anthropogenic impact, was also important. The connection of colour anomalies and human activities shows once
again the importance of amphibians as bioindicators for the early detection of pollution and other harmful effects in aquatic
ecosystems.
Keywords. Abnormal colouration, blue frogs, blue axanthism, leucism, albinism, Ranidae, Salamandridae, Bombinatoridae,
Bufonidae, Human Footprint, pollution, bioindication