[CANCER RESEARCH 38. 210-214. January 1978]
Hydroxylated Metabolites of R,S-1-(Tetrahydro-2-furanyl)-5-
fluorouracil (Ftorafur) in Rats and Rabbits1
A. T. Wu, J. L. Au, and W. Sadee
School of Pharmacy, University of California, San Francisco, California 94143
ABSTRACT
Two hydroxylated metabolites (M and M ) have been
isolated from rabbit urine after administration of Ftorafur
(FT). The structures of 3'-OH-FT and 4'-OH-FT were as
signed to M, and M , respectively. A reverse-phase high
performance liquid chromatography assay was developed
for measuring FT, M,, M . and 5-fluorouracil (FU) plasma
levels. M,, M , and FU were present in rabbit and rat
plasma in greatly varying concentrations after FT admin
istration. Pharmacokinetic studies suggest that FU for
mation proceeds via metabolic intermediate(s) and that
the extent of FT activation is variable. A horse liver
thymidine phosphorylase preparation capable of catalyz
ing the conversion of /¿-ribo-2-deoxy-5-fluorouracN to FU
was inactive against FT and M,. However, 20% of M was
converted to FU by this enzyme, which suggests that the
urinary metabolite M consisted of a mixture of enan-
tiomers with 20% present in the natural /3-D configuration.
The stereochemistry of M, remains unknown. Hydroxyla-
tion of FT to /i-D-4 -OH-FT and subsequent cleavage to
FU by thymidine phosphorylase represents one possible
activation mechanism of FT to FU. However, lack of cor
relation between plasma levels of M and FU indicates
that this mode of metabolic activation may account for
only part of the overall activation of FT in vivo.
INTRODUCTION
The pyrimidine antimetabolite FT3 has shown significant
activity in several adenocarcinomas with a spectrum of
activity similar to but less toxicity than FU (3, 12, 17). FT is
considered to be a chemical depot form of FU (4, 6, 7, 16)
that is generated in vivo, presumably by hepatic metabolism
involving cytochrome P-450 (8, 13). This concept is in
agreement with the rather low in vitro cytotoxic activity of
FT in the absence of significant amounts of cytochrome
P-450 (10). The mechanism of FT conversion to FU remains
unknown.
Several reports support the hypothesis that FU formation
'Supported by USPHS Grant GM-16496 from the National Institutes of
General Medical Sciences and CA-05186 from the National Cancer Institute.
Further support was obtained from Training Grant GM 00728-15; from the
Earl C. Anthony Fund. University of California. San Francisco; and from
Grant RR 00892-01A1 from the Division of Research Resources. NIH, to the
Magnetic Resonance Laboratory, of the University of California, San Fran
cisco.
3To whom requests for reprints should be addressed.
3The abbreviations used are: FT, Ftorafur [1-(tetrahydro-2-furanyl)-5-fluo-
rouracil); FU. 5-fluorouracil; FUdR, /3-ribo-2'-deoxy-5-fluorouracil; GC-MS,
gas chromatography-electron impact-mass spectrometry; HPLC. high-per
formance liquid chromatography; NMR, nuclear magnetic resonance; TLC.
thin-layer chromatography; GC. gas chromatography.
Received June 20. 1977; accepted October 14. 1977.
may not be the only mechanism of FT activation. Smolyan-
skaya and Tugarinov (16) have demonstrated the in vivo
presence of a microbiologically active metabolite fraction
of FT in addition to FU. Differences in FT and FU toxicities
to mice also support an activation mechanism of FT other
than, or in addition to, formation of FU (11), while therapeu
tic cross-resistance between FT and FU suggests that the
major part of the therapeutic mechanism of action is com
mon to FT and FU (9). Pharmacokinetic analysis of FT and
FU plasma levels following administration of FT to rabbits
and rats (18) further indicates the presence of metabolites
other than FU that might be independently active or serve
as intermediate(s) in the formation of FU.
In this study we have used FT specifically labeled either
in the tetrahydrofuranyl ring or in the FU moiety in order to
investigate metabolic pathways of FT other than formation
and subsequent metabolism of FU. With this technique, 2
hydroxylated metabolites of FT were isolated and character
ized, and their presence in circulating peripheral blood of
rats and rabbits was established.
MATERIALS AND METHODS
Chemicals and Reagents. All chemicals and reagents
were of analytical reagent grade. FT was supplied by the
Chemical and Drug Procurement Section, Division of Can
cer Treatment, National Cancer Institute, Washington, D. C.
[2-14C]FT [1-(tetrahydro-2 -furanyl)-5-fluoro-2-14C-uracil;
specific activity, 46 ¿iCi/rng]and [2',5'-14C]FT [1-(tetrahy-
dro-2-furanyl-2',5'-14C)-5-fluorouracil, specific activity, 33
¿iCi/mg]were obtained through the National Cancer Insti
tute from Isotope Synthesis Laboratory, Stanford Research
Institute, Menlo Park, Calif. Bis-15A/-5-fluorouracil (15N,,-FU)
was synthesized from bis-'5/V-thiourea (99% 15N enrich
ment; Koch Isotope, Cambridge, Mass.) (15). /3-Ribo-5-fluo-
rouracil and FUdR were obtained from Nutritional Biochem-
icals Corp., Cleveland, Ohio.
Apparatus. GC was performed on a Varian Aerograph
2700 equipped with a glass column (6 ft x 0.25 inch outside
diameter; 2 mm inside diameter) packed with 3% OV-1 on
Gas-Chrom Q (100 to 120 mesh; Applied Science Laborato
ries, State College, Pa.). The injector, column, and detector
temperatures were 245, 180, and 275°,respectively. Flow
rates for helium, oxygen, and hydrogen were 40, 300, and
30 ml/min, respectively. Mass spectra were obtained by
direct insertion on an AEI MS902 mass spectrometer with
isobutane chemical ionization and by use of a gas Chro
matographie inlet system on a Varian MAT CH-7 electron
impact mass spectrometer connected with a Varian Aero
graph 2700 gas Chromatograph and controlled by a Nova
2-10 computer (GC-MS). 14C activity was measured in a
Searle Analytic Mark III liquid scintillation counter.
210 CANCER RESEARCH VOL. 38
Research.
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