Animal exchange ratios: an alternative point of
view
P. I. D. Kinyua* and J. T. Njoka
Department of Range Management, University of Nairobi, PO Box 29053, Nairobi, Kenya
Abstract
An alternative interpretation is provided of the con-
cepts of carrying capacity and exchange ratios, particu-
larly suitable for game animal species, based on
management models for a given area of rangeland or
pasture. It involves modelling animal population
dynamics as discrete-time logistic equations. Carrying
capacity is then generated endogenously using rainfall
as a proxy. The model interaction parameters, also gen-
erated endogenously, represent the animal exchange
ratios. Because these two parameters are generated
endogenously, this approach takes into account all the
animals' habitat requirements (food, cover, water and
space) simultaneously, unlike other approaches that
tend to consider food requirements only.This makes the
approach amenable to multi-species situations. It also
captures the ecological de¢nition of population growth
models where the realized rather than the theoretical
carrying capacity is determined endogenously.
Key words: carrying capacity, exchange ratios,
population dynamics
Re¨ sume¨
On apporte une interpre¨ tation alternative des concepts
de capacite¨ de charge et de taux d'e¨changes spe¨ ci¢que-
ment adapte¨ e aux espe© ces-gibier, base¨ s sur des mode© les
de gestion pour une zone donne¨ e d'habitat ou de paª tu-
rage. Elle implique la mode¨ lisation de la dynamique des
populations animales en e¨ quations logistiques de¨ limi-
te¨ es dans le temps. La capacite¨ de charge est alors ge¨ n-
e¨re¨ e de manie© re endoge© ne, en utilisant les chutes de
pluie comme proxy. Les parame© tres d'interaction mode¨ -
lise¨s, ge¨ne¨re¨s aussi de manie© re endoge© ne, repre¨sentent
les taux de remplacement des animaux. Comme ces
deux parame© tres sont ge¨ ne¨ re¨ s de cette fac
,
on, cette
approche tient compte en meª me temps de toutes les exi-
gences de l'habitat des animaux (nourriture, couvert
ve¨ge¨ tal, eau et espace), contrairement aux autres
approches qui ont tendance a© ne tenir compte que de la
nourriture. Ceci rend cette approche adaptable a© des
situations touchant de nombreuses espe© ces. Elle inte© gre
aussi la de¨ ¢nition e¨ cologique des mode© les de croissance
de population ou© la capacite¨ de charge e¡ective plutoªt
que the¨ orique est de¨ termine¨ e de manie© re endoge© ne.
Introduction
The development of the concept of exchange ratio and
the related concept of carrying capacity has been con-
centrated on livestock and less so on multi-species,
including game animals (Scarnecchia, 1985; Scarnec-
chia,1990).
Exchange ratio refers to the ratio of the number of
individual animals or animal units of one species or
class of livestock to the number of animals or animal
units of another species or class to achieve a speci¢c
objective, given speci¢c management objective(s), e.g.
multi-species forage harvesting objective (Scarnecchia,
1990). Heady & Child (1994) proposed approximate
exchange ratios based on metabolic body weights that
could be applied as a guide to herbivores with similar
diets, but for animals with di¡erent habitat require-
ments, these exchange ratios should be modi¢ed
according to dietary overlap among species. However,
such modi¢ed exchange ratios should be used with cau-
tion (Scarnecchia, 1985) and have limited practical
application due to complexities of diet selection by her-
bivores. *Correspondence.
#
2001East AfricanWild Life Society, Afr. J. Ecol., 39,59^64 59