Clin Exp Pharmacol Physiol. 2021;48:1059–1069. wileyonlinelibrary.com/journal/cep | 1059 © 2021 John Wiley & Sons Australia, Ltd
Received: 25 December 2020
|
Revised: 6 March 2021
|
Accepted: 21 April 2021
DOI: 10.1111/1440-1681.13510
REVIEW ARTICLE
Spirulina supplementation and oxidative stress and pro-
inflammatory biomarkers: A systematic review and meta-
analysis of controlled clinical trials
Sara Mohiti
1
| Meysam Zarezadeh
2,3
| Fatemeh Naeini
4
| Helda Tutunchi
3
|
Alireza Ostadrahimi
3
| Zohreh Ghoreishi
3
| Mehrangiz Ebrahimi Mamaghani
1
Abbreviations: BDNF, brain-derived neurotrophic factor; BMI, body mass index; COPD, chronic obstructive pulmonary disease; COX-2, cyclooxygenase-2; H
2
O
2
, hydrogen peroxide;
HO, hydroxyl radical; HOO, peroxyl radicals; IkB, inhibitor of nuclear factor kappa B; IL-2, interleukin-2; IL-6, interleukin-6; iNOS, inducible nitric oxide synthase; JNK, c-Jun N-terminal
kinase; MCP-1, monocyte chemoattractant protein--1; MDA, malondialdehyde; NADPH, nicotinamide adenine dinucleotide phosphate hydrogen; NF- κB, nuclear transcription factor κB;
NO, nitric oxide; O
2
, superoxide anion radical; Pc, phytocyanin; p-CREB, p-cAMP response element-binding protein; RNS, reactive nitrogen species; ROS, reactive oxygen species; SMD,
standardized mean difference; TBARS, thiobarbituric acid reactive substances; TNF- α, tumour necrosis factor- α.
1
Nutrition Research Center, Department
of Biochemistry and Diet Therapy, School
of Nutrition and Food Sciences, Tabriz
University of Medical Sciences, Tabriz,
Iran
2
Student Research Committee, Tabriz
University of Medical Sciences, Tabriz,
Iran
3
Nutrition Research Center, Department
of Clinical Nutrition, School of Nutrition
and Food Sciences, Tabriz University of
Medical Sciences, Tabriz, Iran
4
Department of Clinical Nutrition, School
of Nutritional Sciences and Dietetics,
Tehran University of Medical Science,
Tehran, Iran
Correspondence
Meysam Zarezadeh, Nutritional Sciences,
School of Nutrition and Food Sciences,
Tabriz University of Medical Sciences,
Attar-Neishaburi St., Golgasht Alley, Azadi
Blvd., Tabriz, Iran.
Email: meysam.za93@gmail.com
Funding information
Student Research Committee, Tabriz
University of Medical Sciences, Grant/
Award Number: 66477
Abstract
Studies investigating the effects of spirulina on inflammation and oxidative stress
status are controversial. Therefore, the current systematic review and meta-analysis
aimed to evaluate the impacts of spirulina supplementation on oxidative stress indica-
tors and inflammatory markers. PubMed-Medline, SCOPUS, Web of Science, Embase
databases and Google Scholar were searched up to 1 October 2020. Random-effect
analysis was applied to perform meta-analysis. Subgroup analyses and multivariate
meta-regression were performed to find heterogeneity sources. Quality assessment
was conducted using Cochrane Collaboration’s tool. A total of 11 studies that enrolled
465 subjects were included in our meta-analysis. Pooled results demonstrated a sig-
nificant increase in interleukin-2 (IL-2) concentrations [Standardized mean difference
(SMD = 2.69 pg/mL; 95% CI: 0.26, 5.11; P = .03)]; however this result changed to
insignificant (SMD = 0.54 pg/mL; 95% CI: −1.29, 2.27; P > .05) when sensitivity analy-
sis performed. A marginal decreasing effect were also found on interleukin-6 (IL-6)
(SMD = −0.72 mg/dL; 95% CI: −1.50, 0.07; P = .073) and thiobarbituric acid reactive
substances (TBARS) levels (SMD = −0.65; 95% CI: −1.37, 0.08; P = .08). In addition,
results of subgroup analysis revealed a significant reduction in IL-6 and TBARS con-
centrations when the baseline body mass index (BMI) of participants was lower than
25 kg/m
2
. Moreover, spirulina had no significant effect on tumour necrosis factor- α
(TNF- α) (SMD = −0.07 mg/dL; 95% CI: −0.33, 0.18; P = .56) and malondialdehyde
(MDA) concentrations (SMD = −0.42; 95% CI: −0.98, 0.14; P = .14). Spirulina con-
sumption contributed to a significant increase in IL-2 concentrations changing to insig-
nificant after sensitivity analysis and marginal decreasing effects on IL-6 and TBARS
levels. No considerable impacts were observed on TNF- α and MDA concentrations.
KEYWORDS
inflammation, meta-analysis, oxidative stress, spirulina, systematic review