Biochimica et Biophysica A cta, 1176 (1993) 299-304 299
© 1993 Elsevier Science Publishers B.V, All rights reserved 0167-4889/93/$06.00
BBAMCR 13354
Establishment of an enzyme immunoassay system
for gliostatin/platelet-derived endothelial
cell growth factor (PD-ECGF)
Takayoshi Hirano, Kiyofumi Asai, Kohei Matsukawa, Taiji Kato, Masanori Takeuchi,
Masato Yonezawa, Takanobu Otsuka and Nobuo Matsui
Departments of Bioregulation Research and Orthopedic Surgery, Nagoya City University Medical School, Mizuho-ku, Nagoya (Japan)
(Received 12 August 1992)
Key words: Enzyme immunoassay; Gliostatin; Platelet-derived endothelial cell growth factor PD-ECGF
A two-site enzyme immunoassay for gliostatin (GLS)/platelet-derived endothelial cell growth factor (PD-ECGF) has been
developed. The detection limit of gliostatin/PD-ECGF was 30 pg/well, and the optimal assay range was 0.1 to 10 ng/well. This
assay system enabled us to confirm the immunochemical identity of both factors and to detect immunoreactive gliostatin/PD-
ECGF (IR-GLS/PD-ECGF) in human biological body fluids. The age-related analysis from newborn to 69 years revealed that
the serum IR-GLS/PD-ECGF level was high in infants younger than 1 year old (1.8 ng/ml) and in the 20-year-old age group
(1.8 ng/ml), and highest in the umbilical cord blood (2.1 ng/ml). Curiously high concentrations were detected in saliva with a
significant sex difference (11.3 ng/ml for males and 48.7 ng/ml for females), and in synovial fluids (3.7 ng/ml). A number of
human tumor cells, gastric cancer cells, MKN-74, neuroblastoma cells, GOTO, as well as epidermoid carcinoma cells, A431, were
found to produce a significant amount of IR-GLS/PD-ECGF (0.2 to 21.8 ng/mg protein), and some of them secreted the
IR-GLS/PD-ECGF in the conditioned medium (~ 0.5 ng/ml). The enzyme immunoassay system is sufficiently sensitive for the
basic and clinical study of gliostatin/PD-ECGF in human body fluids, tissues and organs.
Introduction
Gliostatin (GLS) is a protein factor, which was ini-
tially identified in neurofibroma type 1 (NF1) extracts
as a growth inhibitory factor against glial cells such as
astrocytes and glial tumor cells, with a relative molecu-
lar mass of 100 kDa comprised of two 50-kDa subunits
[1,2]. This factor has lately been found to play an
essential role on the neuron-glial interaction during
development or regeneration of the nervous system [3].
More recently, we have demonstrated the chemical and
biological identity of gliostatin and platelet-derived en-
dothelial cell growth factor (PD-ECGF) [4], which had
Correspondence to: T. Kato, Department of Bioregulation Research,
Nagoya City University Medical School, Mizuho-ku, Nagoya 467,
Japan.
Abbreviations: GLS, gliostatin; PD-ECGF, platelet-derived endothe-
lial cell growth factor; IR-, immunoreactive-; CM, conditioned
medium; CMF-Tyrode, Ca2+-, Mg2+-free Tyrode's solution; PVDF,
polyvinylidene difluoride; NF1, neurofibromatosis type 1; SDS,
sodium dodecyl sulfate; aFGF, acidic fibroblast growth factor; bFGF,
basic fibroblast growth factor; GMF, glia maturation factor; CNTF,
ciliary neurotrophic factor and PMSF, phenylmethanesulfonyl fluo-
ride.
been purified from platelets and placenta as a growth
factor promoting proliferation and chemotactic migra-
tion of endothelial cells resulting in angiogenesis [5,6].
In this article, we establish a sensitive enzyme im-
munoassay system for gliostatin/PD-ECGF, enabling
the measurement of a minute amount of immuno-
reactive-GLS/PD-ECGF in human biological body flu-
ids.
Materials and Methods
Anti-gliostatin / PD-ECGF antiserum
40 ~g of gliostatin/PD-ECGF, which were purified
from human placenta in accordance with the method
previously described [2,4], were emulsified with 0.2 ml
of complete Freund's adjuvant (Difco, UK) and in-
jected subcutaneously into a New Zealand albino rab-
bit. Booster immunizations of 20 ~g were given after 2
and 5 weeks. Anti-gliostatin/PD-ECGF antibody was
raised 6 weeks after the initial immunization. The
y-globulin fraction of anti gliostatin/PG-ECGF serum
(10 ml) was prepared by two successive steps of ammo-
nium sulfate precipitation (40% and 33% saturation).
The precipitates were collected by centrifugation