Histological evidence of accumulation of iron
in postlarvae of red abalone, Haliotis rufescens
Loreto Andrea Pino Chandia
1
, Andrea C Alfaro
2
, Roberto Flores Aguilar
1
, Marcos Godoy
3
& Alfonso Gutierrez Venegas
1
1
Centro de Investigacio´n y Desarrollo i-mar, Universidad de Los Lagos, Puerto Montt, Chile
2
School of Applied Sciences, Faculty of Health and Environmental Sciences, Auckland University of Technology,
Auckland, New Zealand
3
Panamericana Sur 581, Puerto Montt, Chile
Correspondence: A C Alfaro, Faculty of Health and Environmental Sciences, School of Applied Sciences, Auckland University of
Technology, Private Bag 92006, Auckland 1020 New Zealand. E-mail: andrea.alfaro@aut.ac.nz
Abstract
The effect of iron on abalone postlarvae (Haliotis
rufescens) was investigated in a controlled-cultur-
ing system. Three iron concentrations (0.15, 1.5
and 15 mg L
1
of Fe) and a control (no iron
added) were used to culture H. rufescens postlar-
vae while being fed the diatom Navicula inserta
over 10 days. Results indicate that H. rufescens
postlarvae accumulate iron granules in the stom-
ach, digestive gland and mantle, but not in the
gills or other tissues. The number and diameter
of iron granules in tissues increased with increas-
ing iron concentration in the culturing environ-
ment. The iron accumulation is assumed to have
been acquired in the digestive system through the
iron-enriched diatom feed and in the mantle
through subcutaneous iron transfer. The lack of
iron granules in the gills suggests that iron is not
absorbed through the respiratory system, as is the
case for many filter feeding bivalves. Exposure to
the highest iron concentration (15 mg L
1
)
resulted in tissue abnormalities where granules
accumulated, and may have significantly affected
the health of H. rufescens postlarvae. These find-
ings provide valuable information for the regula-
tion of appropriate iron levels within aquaculture
settings and highlights the importance of moni-
toring iron levels within abalone larval culturing
environments.
Keywords: abalone postlarvae, Haliotis rufes-
cens, iron granules, histological sections, heavy
metal exposure, tissue disintegration
Introduction
For most marine invertebrates, such as abalone,
the larval stage is one of the most vulnerable times
in their life cycle. After the first week in the water
column, abalone larvae undergo initial settlement
and metamorphosis. Once in their benthonic life
stage, postlarvae begin to feed on microalgae, until
they complete the development of the radula and
digestive system to consume macroalgae. The
post-larval period is a critical step in their develop-
ment, and requires a good supply of biofilms with
diatoms and bacteria (Searcy-Bernal 1996). How-
ever, during the first few weeks after settlement,
massive mortalities (>90%) often are observed,
especially if the complex of bacteria and diatom
biofilms are too dense (Hahn 1989; Searcy-Bernal,
Salas-Garza & Flores-Aguilar 1992a; Searcy-Ber-
nal, Salas-Garza, Flores-Aguilar & Hinojosa-Rivera
1992b). This density-dependent mortality appears
to be related to increased susceptibility to post-
larval stresses, such as oxygen saturation and
boundary layer conditions (Searcy-Bernal 1996).
Postlarvae also may be exposed to high concentra-
tions of environmental contaminants (e.g., heavy
metals, pathogens), and chemical fluctuations in
their surroundings, which often cause high mor-
talities. Within these stressful environments, heavy
metals are particularly dangerous for marine
invertebrate larvae and early juveniles, since these
chemicals are easily incorporated and accumulated
in soft tissues (Chang & Reinfelder 2002; Viant,
Walton, TenBrook & Tjeerdema 2002; Ferrer, And-
rade, Asteasuain & Marcovecchio 2006; Gorski &
© 2011 Blackwell Publishing Ltd 1
Aquaculture Research, 2011, 1–8 doi: 10.1111/j.1365-2109.2011.02969.x