This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1111/ced.14113 This article is protected by copyright. All rights reserved DR HUW JOHN MORGAN (Orcid ID : 0000-0002-4542-7572) Article type : Original Article Hair Follicle Differentiation Specific Keratin Expression in Human Basal Cell Carcinoma H. Morgan,* A. Benketah,* C. Olivero, E. Rees, S. Ziaj, A. Mukhtar, S. Lanfredini, G.K. Patel European Cancer Stem Cell Research Institute, School of Biosciences, Cardiff University, Maindy Road, Cardiff, UK Corresponding author: Dr Girish K Patel MD E-mail: patelgk@cardiff.ac.uk * The first two authors contributed equally to this work. Short title: BCC differentiation Abbreviations: Basal cell carcinoma (BCC), Hair follicle (HF), Outer root sheath (ORS), Inner root sheath (IRS), squamous cell carcinoma (SCC). Author contributions: GKP, HM and AB designed research; HM, ER, and AM performed research; HM, SL, SZ, CO and GKP analysed data; and CO, SL, HM, ER, AM and GKP wrote the manuscript. Conflict of interest: None declared. Funding: Ministry of Higher Education and Scientific Research (Libya), Cancer Research UK. Accepted Article