Pharmacology Biochemistry and Behavior, Vol. 42, pp. 509-515, 1992 0091-3057/92 $5.00 + .00
Printed in the U.S.A. All rights reserved. Copyright © 1992 Pergamon Press Ltd.
Cocaine In Utero Enhances the Behavioral
Response to Cocaine in Adult Rats
JOANNA PERIS, l
MIA COLEMAN-HARDEE AND WILLIAM J. MILLARD
University of Florida, Gainesville, FL 32610
Received 16 December 1991
PERIS, J., M. COLEMAN-HARDEE AND W. J. MILLARD. Cocaine in utero enhances the behavioral response to
cocaine in adult rats. PHARMACOL BIOCHEM BEHAV 42(3) 509-515, 1992.--The effects of cocaine exposure in utero
on cocaine-induced behaviors and dopamine (DA) transmission in the nigrostriatal and mesolimbic pathways were measured
in adult rats. Pregnant rats received either saline or cocaine (1 or 3 mg/kg, IV) daily throughout gestation. When offspring
were 3 months of age, locomotor and stereotypic behaviors were rated after an injection of either saline or cocaine (10 mg/
kg, IP). Cocaine in utero increased the response to cocaine in adult offspring and increased basal locomotion in female
offspring. Cocaine in utero increased amphetamine-stimulated release in female offspring but decreased release in males. On
the other hand, male rats that had received cocaine in utero exhibited greater basal tritium release. One injection of cocaine
increased amphetamine-stimulated [3H]DA release from striatal slices of male rats but not female rats. Neither cocaine in
utero nor in vivo affected D2 DA receptor binding in striatum nor nucleus accumbens. Thus, cocaine in utero behaviorally
sensitized animals to subsequent cocaine exposure and increased [3H]DArelease from nigrostriatal endings, but the relation-
ship of these two variables depended upon gender.
Cocaine In utero Dopamine Striatum Stereotypy Locomotion
COCAINE use has increased significantly in young women in
their child-bearing years and unfortunately as many as 17070
of these women continue to use cocaine during pregnancy
(14,25). The effects of cocaine use during pregnancy include
shorter gestationai periods and offspring with decreased birth
weights, increased incidence of urogenital malformations, and
observable behavioral deficits (5-7,29). These findings have
been reproduced in laboratory animals such that fetal expo-
sure of at least 60 mg/kg/day decreases birth weight and in-
creases physical malformations (8,13,26) while lower doses
decrease learned behaviors and increase spontaneous locomo-
tor activity up to 30 days postnataily (15,38,39). Similarly,
exposure to cocaine in rats during postnatal days 1-10 (a time
period comparable to third trimester exposure in humans) re-
sults in a slight increase in basal locomotion but a twofold
increase in amphetamine-induced locomotion (20). Thus, in
utero exposure to low doses of cocaine appears to increase
locomotor activity, especially in response to another stimu-
lant, and this effect may persist into adulthood. These data
suggest that cocaine in utero may affect the adult response to
cocaine although this has not yet been measured.
Cocaine exposure in adult rats increases the degree of loco-
motion and stereotypy induced by subsequent cocaine admin-
istration and this sensitization is long lasting [see (41)]. The
mesolimbic and nigrostriatal dopaminergic pathways in the
brain are thought to mediate the behavioral effects of cocaine
(9) and increased dopamine (DA) transmission in these path-
ways has been proposed as a mechanism for cocaine sensitiza-
tion. More specifically, cocaine sensitization results in en-
hanced striatai DA release (1,22,32-35) and postsynaptic D~
and D2 DA receptor responsiveness in nucleus accumbens
(16,17,32). If the behavior response to cocaine in adult rats
is sensitized by in utero exposure to cocaine, then it is also
possible that DA transmission in nigrostriatai or mesolim-
bic pathways may be enhanced after cocaine exposure in
utero.
The purpose of the work reported here was to test whether
in utero cocaine can result in sensitization either behaviorally
or neurochemicaily in adult offspring. Specifically, we asked
whether the initial behavioral response to cocaine as an adult
was affected by in utero exposure to cocaine and whether
nigrostriatal and mesolimbic DA terminals were affected by
in utero exposure to cocaine. We measured locomotor activity
and stereotypy, [3H]DA release from striatum, and D2 DA
receptor binding in striatum and nucleus accumbens both after
a saline or cocaine in vivo challenge in adult rats that had
been treated in utero with either saline or one of two doses of
cocaine.
Requests for reprints should be addressed to Joanna Peris, Ph.D., Box 100487, J. Hillis Miller Health Center, University of Florida,
Gainesville, FL 32610.
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