Synonymy of Philippine Ixora (Ixoreae, Rubiaceace) and
Ixora silagoensis sp. nov.
Cecilia I. Banag, Nicole Marie B. Manalastas and Grecebio Jonathan D. Alejandro
C. I. Banag (cibanag@mnl.ust.edu.ph), N. M. B. Manalastas and G. J. D. Alejandro, Dept of Biological Sciences, College of Science, Univ. of
Santo Tomas España, Manila 1015, Philippines. CIB and GJDA also at: Research Center for the Natural and Applied Sciences and Graduate
School, Univ. of Santo Tomas España, Manila 1015, Philippines.
Two endemic Philippine species of Ixora collected in southern Leyte were studied, and investigation of type and other
herbarium specimens revealed that one of the species is I. longissima Merr., and that the other does not match any of
the known Philippine Ixora species. e recollection of I. longissima after almost a century was essential to investigate
whether or not it is conspecific with I. salicifolia DC. is study supports the synonymy of I. longissima and I. salicifolia
and provides a full description and illustration of the new species, Ixora silagoensis Manalastas, Banag & Alejandro.
e conservation status of I. silagoensis is assessed as ‘Critically Endangered’ (CR) based on the criteria of IUCN.
Ixora L. (Ixoreae), is the third largest genus of the family
Rubiaceae with ca 530 species ranging from shrubs to small
trees (Mouly and Pisivin 2007). Due to the large number of
species in the genus, it is presently subject to important
investigations and taxonomic changes (Mouly 2007). In the
recently circumscribed tribe Ixoreae, Ixora was further
enlarged to accommodate several genera including
Captaincookia N. Hallé, Doricera Verdc., Hitoa Nadeaud,
Myonima Comm. ex Juss., Sideroxyloides Jacq., ouarsiora
Homolle ex Arénes, and Versteegia Valeton (Mouly et al.
2009b). Despite some previous and on-going studies on
Ixora, the continental Asian, American and Pacific taxa
have never been fully revised and no clear picture of their
diversity is available (De Block 1998, Mouly 2007, Mouly
et al. 2009a, b). According to Merrill (1923), about 42
species of Ixora have been recorded in the Philippines, 32 of
which are endemic. is account was updated by Alejandro
(2007), identifying 30 Philippine endemic Ixora species out
of 41 species. No other studies on Philippine Ixora have been
done since then. us, the Ixora species of the Philippines
still lack comprehensive descriptions, illustrations and taxo-
nomic keys. Information on their distribution, ecology and
conservation status is also limited to non-existent.
While studying the protologues and examining speci-
mens of Ixora for a revision of the Philippine species, several
names that needs to be updated came to light. Some
collections, on the other hand, could not be matched with
known species from the Philippines or neighboring coun-
tries and most likely represent new species that need to be
described. Among them are the two species collected in
Silago and Sogod of the Leyte province. e species collected
in Sogod was identified as Ixora longissima Merr., recollected
after almost a century since its last collection in 1913. Ixora
longissima is endemic to Leyte and was described by Merrill
(1910) as closely allied with I. salicifolia (Blume) DC., a
non-endemic species of Ixora in the Philippines also found
in Indonesia and Malaysia. Both species are characterized
by very long leaves. Due to lack of material and published
literature on Philippine Ixora, the synonymy of these
two species was neither questioned nor investigated. is
study is the first attempt to study whether I. longissima
and I. salicifolia are the same species and to describe an
Ixora species collected in Silago which differ from all other
Philippine Ixora species in terms of its inflorescence.
Material and methods
Taxon sampling
is study was based on field observations of two species of
Ixora collected in the province of Leyte and examination of
herbarium specimens of Ixora from the following herbaria:
BR, C, HUH, K, NY, P, PNH, US. Fieldwork were con-
ducted in Barangay Catmon, Silago, Leyte and Barangay
San Juan, Sogod, and Leyte. Vouchers are deposited at
USTH. e herbarium material used for the morphological
study of I. salicifolia and I. longissima is listed in Table 1.
Preliminary observations on the morphological features
of the Ixora species were noted in the field during collection.
Further examination of the vegetative and reproductive
parts was done on preserved collections using an Olympus
Nordic Journal of Botany 32: 761–764, 2014
doi: 10.1111/njb.00411
© 2014 e Authors. Nordic Journal of Botany © 2014 Nordic Society Oikos
Subject Editor: John Parnell. Accepted 25 October 2013
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