Received: 14 December 2022 Accepted: 20 April 2023
DOI: 10.1002/bab.2469
REVIEW ARTICLE
An insight into microbial sources, classification, and
industrial applications of xylanases: A rapid review
Damanjeet Kaur
1
Amit Joshi
1
Varruchi Sharma
1
Navneet Batra
2
Anil K. Sharma
3
1
Department of Biotechnology, Sri Guru
Gobind Singh College, Chandigarh, India
2
Department of Biotechnology, GGDSD
College, Chandigarh, India
3
Department of Biotechnology, Maharishi
Markandeshwar Engineering College,
Maharishi Markandeshwar (deemed to be
University), Mullana-Ambala, Haryana,
India
Correspondence
Navneet Batra, Department of
Biotechnology, GGDSD College, Sector 32,
Chandigarh 160032, India.
Email: navneet05@gmail.com
Anil K. Sharma, Department of
Biotechnology, Maharishi
Markandeshwar Engineering College,
Maharishi Markandeshwar (deemed to be
University), Mullana-Ambala 133207,
Haryana, India.
Email: anibiotech18@gmail.com
Abstract
Endo 1,4-β-d-xylanases (EC3.2.1.8) are one of the key lignocellulose hydrolyz-
ing enzymes. Xylan, which is present in copious amounts on earth, forms
the primary substrate of endo-xylanases, which can unchain the constituent
monosaccharides linked via β-1,4-glycosidic bonds from the xylan backbone.
Researchers have shown keen interest in the xylanases belonging to glycoside
hydrolase families 10 and 11, whereas those placed in other glycoside hydro-
lase families are yet to be investigated. Various microbes such as bacteria and
fungi harbor these enzymes for the metabolism of their lignocellulose fibers.
These microbes can be used as miniature biofactories of xylanase enzymes for a
plethora of environmentally benign applications in pulp and paper industry, bio-
fuel production, and for improving the quality of food in bread baking and fruit
juice industry. This review highlights the potential of microbes in production of
xylanase for industrial biotechnology.
KEYWORDS
hemicellulose, lignocellulosic biomass, xylanases, xylolytic enzymes
1 INTRODUCTION
Hemicellulose, lignin, and cellulose are the principal lig-
nocellulosic biomass in the cell wall of plants, which
are associated with each other by both the covalent and
noncovalent bonds. Further, 25%–30% of all the lignocel-
lulose of plants is made up of hemicellulose.
1
Besides
this, hemicellulose is itself composed of complex het-
eropolysaccharide, that is xylan.
2
Xylan is a polymer of
β-xylopyranose residues, which are repeatedly joined to
each other with the help of β-1,4-glycosidic bonds.
3
These
β-xylopyranose residues are further substituted with sim-
Abbreviations: AP, alkaline phosphate; SOD, superoxide dismutase.
Damanjeet Kaur and Amit Joshi contributed equally to this work.
© 2023 International Union of Biochemistry and Molecular Biology, Inc.
ple sugars and organic acids.
4
Further, the importance of
xylan can be deduced from the fact that it ranks second
after cellulose in terms of its global abundance.
5
It occurs
naturally both in hardwoods and softwoods. Hence, it can
be employed for several biotechnological applications by
enzymatically converting it into simpler forms of sugars.
6–8
Although complete depolymerization of xylan requires an
alliance of a set of xylolytic enzymes like endo 1,4-β-d-
xylanases (EC3.2.1.8), which play a pivotal role in cleaving
xylan specifically at 1,4-β-gylcosidic bonds connecting the
primary chain of xylopyranose residues. Xylanases belong-
ing to glycoside hydrolase (GH) families 10 and 11 have
been explored extensively more than those belonging to
GH familes 5, 7, 8, and 43.
9,10
Biotechnol Appl Biochem. 2023;1–15. wileyonlinelibrary.com/journal/bab 1