International Journal of Plant Research 2016, 6(3): 64-87
DOI: 10.5923/j.plant.20160603.03
Effect of Gibberellic Acid, Paclobutrazol and Zinc on
Growth, Physiological Attributes and the Antioxidant
Defense System of Soybean (Glycine max) under Salinity
Stress
Mahmoud R. Sofy
Botany and Microbiology Department, Faculty of Science, Al-Azhar University, 11884 Nasr City, Cairo, Egypt
Abstract Soil salinity is one of the major abiotic stresses which caused significant reduction in the growth parameters,
photosynthetic pigments and yield components of soybean plants. The present study aims to improvement of soybean
production under saline conditions and also tries to elucidate the possible mechanisms of plant tolerance by using three
different treatments (Gibberellic acid, Paclobutrazol and Zinc sulphate). The magnitude of reduction, increased by increasing
salinity level. Application of the above mentioned treatments, in the absence and presence of NaCl, has greater changes in
most of the assayed parameters where the adverse effects of salinity as regards the growth characters, photosynthetic
pigments and yield components as well as soluble carbohydrates, soluble protein and oil contents in the yielded seeds were
significantly mitigated by treatment with either GA
3
, PBZ or Zn. Salinity caused significant decreases in the activities of
endogenous gibberellic acid (GA
3
) and indole acetic acid (IAA) while activities of both Jasmonic acid (JA) and Abscisic acid
(ABA) were increased. Significant increases were observed in the activities of superoxide dismutase (SOD), peroxidase
(POX), catalase (CAT) and glutathione reductase (GR) in shoots of salt stressed plants. Application of GA
3
, PBZ or Zn
caused great variations in the activities of endogenous phytohormones and antioxidant enzymes. Treatment with either GA
3
,
PBZ or Zn caused significant reduction in both lipid peroxidation and proline that observed in shoots and root of salinized
soybean plants. The electrophoregram of protein pattern of the yielded seeds of the soybean in response to their
pre-emergence treatment with different concentrations of NaCl (50 mM & 100 mM) with or without GA
3
, PBZ or Zn
appeared variations in the number of electrophoretic protein bands. A total of 23 bands was detected with different molecular
weights ranging from 129 kDa to 9 KDa. Two newly formed protein bands have appeared, 1 of them (16 KDa) at 50 mM with
Zn or PBZ and the other band (19 KDa) has appeared at 50 mM NaCl with PBZ. Generally, it could be concluded that
application of either GA
3
, PBZ or Zn have (to more extent) a beneficial regulatory role in plants grown under salt stress
conditions.
Keywords Gibberellic acid, Paclobutrazol, Zinc, Antioxidant defense system, Soybean (Glycine max), Salinity stress
1. Introduction
Worldwide, 20% of total cultivated and 33% of irrigated
agricultural lands are exacerbated by high salinity.
Phenomena like low precipitation, high surface evaporation,
irrigation with saline water, weathering of native rocks, and
poor agricultural practices have increased the rate of soil
salinization to 10% per annum. It has been predicted that
more than 50% of the arable land would be salinized by the
year 2050 [1].
High salt levels generate a two-component stress on plants:
* Corresponding author:
mahmoud_sofy@yahoo.com (Mahmoud R. Sofy)
Published online at http://journal.sapub.org/plant
Copyright © 2016 Scientific & Academic Publishing. All Rights Reserved
an osmotic stress caused by reducing water availability in
soil and an ionic stress due to imbalance of solutes in the
cytosol [2]. Recent research has identified various adaptive
responses to salinity stress at molecular, cellular, metabolic,
and physiological levels, although mechanisms underlying
salinity tolerance are far from being completely understood.
This paper provides a comprehensive review of major
research advances on biochemical, physiological, and
molecular mechanisms regulating plant adaptation and
tolerance to salinity stress [3].
Soybean (Glycine max L.) a legume species native to East
Asia, is now widely grown as the primary oilseed crop in the
world including in many regions in the world. It is
noteworthy that China is currently the largest importing
country for soybean despite being one of its origins. To meet
the increasing demand for food, oil and protein resources,