Citation: Antúnez, P.; Hernández-
Cruz, I.; Ibrahim-Abdulsalam, F.;
Clark-Tapia, R.; Ruiz-Aquino, F.;
Valenzuela-Encinas, C. Effects of
Distance and Neighbor Size on
Abies hickelii: The Asymmetric
Competition Is Aggravated in an
Endangered Species. Forests 2023, 14,
1654. https://doi.org/10.3390/
f14081654
Academic Editor: Andrew M. Barton
Received: 30 June 2023
Revised: 12 August 2023
Accepted: 13 August 2023
Published: 16 August 2023
Copyright: © 2023 by the authors.
Licensee MDPI, Basel, Switzerland.
This article is an open access article
distributed under the terms and
conditions of the Creative Commons
Attribution (CC BY) license (https://
creativecommons.org/licenses/by/
4.0/).
Article
Effects of Distance and Neighbor Size on Abies hickelii: The
Asymmetric Competition Is Aggravated in an Endangered Species
Pablo Antúnez
1,
* , Iván Hernández-Cruz
2
, Fatima Ibrahim-Abdulsalam
3
, Ricardo Clark-Tapia
1
,
Faustino Ruiz-Aquino
1
and César Valenzuela-Encinas
1
1
División de Estudios de Posgrado, Instituto de Estudios Ambientales, Universidad de la Sierra Juárez,
Avenida Universidad S/N, Ixtlán de Juárez, Oaxaca 68725, Mexico
2
Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Avenida Universidad S/N, Ixtlán de Juárez,
Oaxaca 68725, Mexico; ivanhernandezcruz10@gmail.com
3
School of Public Health, Walailak University, Thasala 80160, Thailand
* Correspondence: pantune@uni-goettingen.de; Tel.: +55-951-5536362
Abstract: One of the hypotheses in the field of forest ecology suggests that competition among
individual plants is asymmetric or unidirectional. This notion holds particular significance for
endangered species, necessitating a comprehensive assessment of the impact of their closest com-
petitors to understand their spatial distribution and local-scale interactions. The mean response of
Abies hickelii’s diameter at breast height and total height was determined for each unit that increased
the distance from its neighbor and each unit that changed the size of its neighbor. The results obtained
through the analysis of covariance suggest that, for each meter that the distance from the neighbor
increases, there is a mean gain of 1.13 cm in the diameter of the individuals studied. The height results
showed higher sensitivity to changes in the neighbor distance than the diameter. It was observed
that the population is exposed to asymmetric competition, suggesting that larger individuals take
available resources disproportionately, thereby inhibiting the growth of smaller individuals. The
intraspecific competition surpassed the interspecific competition. A suggestion was made to use
selective thinning during the early-growth stages of the species. The habitat of the species studied is
under strong pressure from land use change due to clandestine logging, the opening of roads, and
other anthropogenic effects. Thus, these results represent an important step in the implementation of
action strategies to preserve the species in the wild.
Keywords: Cohen’s index; effect of first five neighbors; endangered species; Mexican fir; Weibull
1. Introduction
Abies hickelii (Flous et Gaussen) is a critically endangered species found in isolated
populations forming strips or small patches in the mountain ranges of Mexico, mainly in
Chiapas, Oaxaca, and Veracruz [1]. Currently, it is listed in the endangered category on the
IUCN Red List, and its population is decreasing [2,3]. This species has also been recognized
as a priority for conservation in the official Mexican standard NOM-059-SEMARNAT-2010
related to the conservation of the diversity of species in wildlife (NOM-059, 2010). Small
and isolated populations like A. hickelii are particularly vulnerable to competition for lim-
ited resources, which can be exacerbated by their low genetic diversity and increased
inbreeding [4,5]. These factors hinder the species’ adaptive capacity and restrict its
ability to expand its territory or thrive in the wild. Considering the context of climate
change and land use alterations, understanding the factors influencing the growth and
survival of geographically restricted populations like A. hickelii becomes vital for effective
conservation efforts [6,7].
Understanding the pattern of spatial distribution of plants in wildlife in accurate
and meaningful terms has not been an easy task due to various factors, including
interactions such as competition and interference [8]. Conventionally, the effect of
Forests 2023, 14, 1654. https://doi.org/10.3390/f14081654 https://www.mdpi.com/journal/forests