Flexibly encoded GWAS identifies novel nonadditive SNPs in individuals of African and European ancestry Jiayan Zhou 1,2 , Lindsay Guare 3 , Andre Luis Garao Rico 1 , Tomas Gonzalez Zarzar 1 , Nicole Palmiero 1 , Themistocles L. Assimes 2 , Shefali Setia Verma 4 , Molly Ann Hall 1,5,6 1. Department of Veterinary and Biomedical Sciences, College of Agricultural Sciences, The Pennsylvania State University, University Park, PA 16802, USA 2. Division of Cardiovascular Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, CA 94304, USA 3. Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA 4. Department of Pathology and Laboratory Medicine, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA 5. The Huck Institutes of the Life Science, The Pennsylvania State University, University Park, PA 16802, USA 6. Penn State Cancer Institute, The Pennsylvania State University, University Park, PA 16802, USA All rights reserved. No reuse allowed without permission. (which was not certified by peer review) is the author/funder, who has granted medRxiv a license to display the preprint in perpetuity. The copyright holder for this preprint this version posted June 5, 2023. ; https://doi.org/10.1101/2023.06.01.23290857 doi: medRxiv preprint NOTE: This preprint reports new research that has not been certified by peer review and should not be used to guide clinical practice.