1022-7954/03/3906- $25.00 © 2003 MAIK “Nauka /Interperiodica” 0700
Russian Journal of Genetics, Vol. 39, No. 6, 2003, pp. 700–705. Translated from Genetika, Vol. 39, No. 6, 2003, pp. 840–846.
Original Russian Text Copyright © 2003 by Marussin, Puzyrev, Salyukov, Bragina.
INTRODUCTION
Oxidative stress, i.e., an increase in formation of
active oxygen forms and lipid peroxides (LPs), plays an
important role in pathogenesis of a wide range of dis-
eases [1–5]. The multicomponent, integrated and regu-
lated antioxidant system (AOS) prevents uncontrolled
free-radical oxidation (FRO) and maintains the optimal
balance of redox reactions [6–11]. At present, the con-
tribution of genetic factors into variation of the AOS
parameters is studied. This may be related to the fact
that the entire complex of interacting and interchange-
able low- and high-molecular substances is difficult to
analyze; moreover, the technical approaches used are
often inadequate to assess the state of AOS. By using a
single model system, it is impossible to assess unam-
biguously and comprehensively all components of the
antioxidant defense, such as scavenging free radicals,
hydrogen and lipid peroxide degradation and restora-
tion, binding metal ions that undergo valence changes
and serve as potential catalytic agents for FRO, biomol-
ecule repair after oxidative damage, and utilization of
FRO products [12–14]. According to the literature data
[15] and the results of our studies [16], about 50% of
variation of the blood plasma antioxidant activity
(AOA) is accounted for by additive effects of many
genes. In view of that, the search for and specification
of the AOS genetic determinants are of importance.
We have studied the contribution of genetic varia-
tion of transferrin (TF) and angiotensin-converting
enzyme (ACE) to the population variation of the blood
plasma AOA determined in the Fe
2+
–lecithin model
system from the yield of the products interacting with
thiobarbituric acid (TBA).
The TF antioxidant effect results from binding iron
ions. The AOA decreases with apoenzyme saturation
with iron, though the complete saturation may lead,
conversely, to generation of free radicals inducing the
peroxide effect [6]. ACE does not belong to the antiox-
idant enzymes. Nevertheless, its polymorphism may be
associated with AOA variation, because of multiple
physiological and biochemical ACE functions, the
major of which is blood pressure regulation [17]. ACE
regulates the content of angiotensin II, which increases
the production of superoxide anions through
NADP/NADPH-dependent oxidases [18, 19]. Of inter-
est is the association between the insertion/deletion
(I/D) polymorphism of the ACE gene (in the Alu repeat)
and the content and activity of the enzyme in blood
plasma [20, 21].
MATERIALS AND METHODS
The blood plasma AOA was determined in the unre-
lated subjects inhabiting the city of Seversk (75 males
and 46 females) and Kargala village (38 males and
40 females) of the Tomsk oblast. The average age of
the subjects examined was 52.88 ± 0.89 and 33.14 ±
1.29 years in the urban and rural populations, respec-
tively. The samples of heparinized venous blood were
taken during a period from October 1996 to June 1998.
In the urban population, genotyping for the TF and
ACE loci was conducted in 130 and 141 subjects,
respectively, of which 102 and 111 subjects were exam-
Correlation between Polymorphism
of the Genes for Transferrin and Angiotensin-Converting
Enzyme and Antioxidant Activity of Blood Plasma
A. V. Marussin, V. P. Puzyrev, V. B. Salyukov, and E. Yu. Bragina
Research Institute of Medical Genetics, Tomsk Research Center, Russian Academy of Medical Sciences, Tomsk, 634050 Russia;
fax: (3822) 51-37-44; e-mail: marussin@img.tsu.ru
Received March 13, 2002; in final form June 13, 2002
Abstract—In 75 male and 46 female subjects of an urban population (93% Russians) and in 38 males and
40 females of a rural population (87% Russians), the antioxidant activity (AOA) of blood plasma was deter-
mined from the plasma ability to reduce the yield of products interacting with thiobarbituric acid in the model
lecithin–Fe
2+
ion system. In the urban population, the loci TF (AvaI in exon5) and ACE (I/D polymorphism of
the Alu repeat in intron16) were studied in 130 and 141 subjects, respectively. Of them, 102 and 111 subjects,
respectively, were examined for AOA. In the rural population, the corresponding sample sizes were 75 and 76
(73 and 74 subjects were examined for AOA). The polymorphic loci of the urban and rural populations did not
differ in the allele frequencies. In both populations Hardy–Weinberg and gametic equilibria were observed. The
contributions of the TF and ACE genes to AOA variation in the combined sample from the urban and rural pop-
ulations were 0.6 and 0.5%, respectively.
HUMAN
GENETICS