Epidemiologic Reviews
Copyright © 1996 by The Johns Hopkins University School of Hygiene and Public Health
All rights reserved
Vol. 18, No. 1
Printed in U.S.A.
Developing an Integrated Epidemiologic Approach to Emerging Infectious
Diseases
Stephen S. Morse
1
and James M. Hughes
2
Epidemiology has been a major contributor to the
success of the disease control efforts of the past cen-
tury, culminating in such signal triumphs as the global
eradication of smallpox and the eradication of polio
from the Western Hemisphere. However, in recent
years, partly because these very successes led to a
pervasive optimism about infectious diseases in the
future, there has been a waning interest in infectious
disease epidemiology even though infectious diseases
remain the leading cause of death worldwide and an
important cause of death in the United States (1-3). In
addition, the tragic recent explosion of the acquired
immunodeficiency syndrome (AIDS) pandemic and
the emergence of multiple drug resistant tuberculosis
forcefully reminded us that infectious diseases could
not be relegated to the past.
AIDS, like many of the plagues of the past, falls into
the category of emerging infections, seemingly new
diseases that appear suddenly and unexpectedly.
Emerging infections can be defined as those that either
have newly appeared in a population or that are rapidly
increasing their incidence or expanding their geo-
graphic range (4, 5). Other recent examples include
hantavirus pulmonary syndrome, Lyme disease, hem-
orrhagic colitis, and hemolytic uremic syndrome (re-
sulting from a foodborne infection caused by certain
strains of Escherichia coli), and Ebola hemorrhagic
fever in Africa (6). Past scourges can also recur and
are referred to as reemerging diseases, which are often
conventionally understood and well recognized public
health threats that have increased or reappeared be-
cause previously active public health measures have
lapsed or sanitary infrastructure has deteriorated. Spe-
cific factors precipitating disease emergence can be
identified in virtually all cases (2, 4, 5). These include
Received and accepted for publication February 20, 1996.
Abbreviations: AIDS, acquired immunodeficiency syndrome; HIV,
human immunodeficiency virus.
1
Columbia University, School of Public Health, New York, NY.
2
Director, National Center for Infectious Diseases, Centers for
Disease Control and Prevention, Atlanta, GA.
Reprint requests to Dr. Stephen S. Morse, Division of Epidemi-
ology, Columbia University School of Public Health, 600 West 168th
Street (PH-18), New York, NY 10032.
ecological or environmental factors that place people
in increased contact with a pathogen or its natural host,
and environmental, social, demographic, and behav-
ioral factors that promote establishment and dissemi-
nation of a pathogen previously introduced into a
smaller population.
The conditions of modern life and changes in human
behaviors make these factors more prevalent, giving
reason to expect emerging diseases to increase. His-
torically, "new" diseases have appeared and spread as
by-products of exploration, trade, or warfare, when the
movement of people, animals, or goods brought geo-
graphically isolated infections to new grounds (7). In
the nineteenth century, steamships carried cholera to
Europe and Africa. Today, trucks, freighters, and air-
planes have largely replaced caravans and steamships,
allowing even richer opportunities for infections to
emerge and spread efficiently. Speed of travel and
global reach of infections are borne out by studies
modeling the spread of influenza epidemics (8) and of
human immunodeficiency virus (HIV) (9, 10), as well
as by the actual progress of known epidemics.
Other factors are also allowing emerging infections
to appear at increasing rates and could facilitate wider
and more rapid spread. In many parts of the world,
economic conditions are encouraging the mass move-
ment of workers from rural areas to cities. It has been
estimated that, largely as a result of this migration, by
the year 2025, 65 percent of the world population,
including 61 percent of the population in developing
regions, will live in cities (11). The phenomenon of
rural to urban migration can allow infections arising in
isolated rural areas, which may once have remained
unrecognized and localized, to reach larger popula-
tions, with the city serving an amplifying function. An
infection can become further disseminated when other
migrants return home with an infection acquired in the
city, a pattern now being observed with HIV in Asia.
The history of HIV, in fact, is a case in point. HIV
may have originated as a zoonotic introduction in a
place where people were in contact with the natural
host (12). After its probable first move from a rural
area into a city, HIV-1 spread along highways to other
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