Reprod Dom Anim. 2018;1–6. wileyonlinelibrary.com/journal/rda
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1 © 2018 Blackwell Verlag GmbH
1 | INTRODUCTION
Disorders of sex development (DSDs) are responsible for sterility
or infertility among domestic animals (Szczerbal & Switonski, 2016).
These disorders can be caused by gene mutations or chromosome
anomalies, as well as by other factors, such as placental anastomosis
of heterosexual fetuses. The nomenclature of DSDs, initially devel‐
oped for humans (Hughes, Houk, Ahmed, & Lee, 2006) and adopted
for dogs and cats (Poth, Breuer, Walter, Hecht, & Hermanns, 2010),
is based on sex chromosome status related to gonadal and pheno‐
typic sex. Analysis of these chromosomes should therefore be the
first step in diagnosis.
A normal male sex chromosome complement (38,XY) has very
often been observed in DSD cats (Nowacka‐Woszuk et al., 2014;
Pienkowska‐Schelling, Becker, Pineroli, & Schelling, 2015; Reynolds
et al., 2014; Schlafer et al., 2011). On the contrary, testicular XX
Received: 22 March 2018
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Revised: 18 May 2018
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Accepted: 3 June 2018
DOI: 10.1111/rda.13263
ORIGINAL ARTICLE
Disorders of sex development in cats with different
complements of sex chromosomes
Izabela Szczerbal
1
| Paulina Krzeminska
1
| Stanislaw Dzimira
2
| Tuire Maria
Tamminen
3
| Seppo Saari
4
| Wojciech Nizanski
5
| Maciej Gogulski
6
| Joanna Nowacka‐
Woszuk
1
| Marek Switonski
1
1
Department of Genetics and Animal
Breeding, Poznan University of Life Sciences,
Poznan, Poland
2
Department of Pathology, Wroclaw
University of Environmental and Life
Sciences, Wroclaw, Poland
3
Department of Production Animal
Medicine, University of Helsinki, Helsinki,
Finland
4
Veterinary Histopathology Service Patovet
Ay Kivihaantie 7, Helsinki, Finland
5
Department of Reproduction and Clinic
of Farm Animals, Wroclaw University of
Environmental and Life Sciences, Wroclaw,
Poland
6
University Centre for Veterinary
Medicine, Poznan University of Life
Sciences, Poznan, Poland
Correspondence
Marek Switonski, Department of Genetics
and Animal Breeding, Poznan University of
Life Sciences, Poznan, Poland.
Email: switonsk@up.poznan.pl
Funding information
Statutory fund of the Department of
Genetics and Animal Breeding, Poznan
University of Life Sciences, Poland, Grant/
Award Number: 508.534.00.0
Abstract
The genetic background of disorders of sex development (DSDs) in cats is poorly
understood, due to a relatively low number of such studies in this species. Here we
present three new DSD cases with different complements of sex chromosomes. The
first, an Oriental Shorthair cat with a rudimentary penis, abdominal atrophic testicles
and lack of uterus appeared to be a freemartin, since leucocyte chimerism XX/XY
and a lack of Y‐linked genes (SRY and ZFY ) were observed in DNA isolated from hair
follicles. XXY trisomy was identified in the second case, a tortoiseshell Devon Rex
male cat with atrophic scrotal testicles and a normal penis. Finally, a European
Shorthair cat with atrophic testicles in a bifid scrotum, rudimentary penis and a lack
of uterus had XY complement, including Y chromosome of normal size and morphol‐
ogy. Also presence of eight Y‐linked genes, detected by PCR, was confirmed. Due to
the low testosterone level in this last patient, we searched for a causative mutation
in two candidate genes (HSD3B2 and HSD17B3) involved in the metabolism of this
steroid hormone. Altogether, five polymorphic sites in HSD3B2 and two in HSD17B3
were found, but none of them showed associations with DSD phenotype. We thus
excluded a possibility that the causative mutation is present in these genes. In con‐
clusion, we confirmed that analysis of the sex chromosome complement is a crucial
step in diagnosis of DSDs. However, extensive molecular studies of the genes in‐
volved in sex development are needed to elucidate the causes of DSDs in cats with
normal complements of sex chromosomes.
KEYWORDS
aneuploidy, intersexuality, leucocyte chimerism, XXY trisomy