Calcium Quantitation with a Parallel Electron Energy Loss
Spectroscopy/Cooled Charge-Coupled Device/200
keV System
Ruoya Ho, Jianglin Feng, Zhifeng Shao, and Andrew P. Somlyo*
Department of Molecular Physiology and Biological Physics, University of Virginia, HSC, Box 449, Charlottesville, VA 22908
Abstract: We evaluated factors affected the accuracy and precision of quantitating trace concentration of Ca
with electron energy loss spectroscopy (EELS). These factors include internal reflection in the spectrometer,
precision of correlation between standards and experimental spectra, and radiation damage–induced spectral
changes. We present methods of correcting for these effects and improving the reliability of trace Ca quanti-
tation. A two-step fitting procedure is described that improves the retrieval of small Ca signals from the large
background common to biological specimens. After optimizing the experimental conditions and data process-
ing procedures, our current system can detect about 2.2 mmol/kg Ca in a 730-Å thick specimen at a total dose
of about 410 nA ? sec at 95% confidence level by fitting the first difference spectra. Because of the 0.1% residual
gain variation after gain normalization, the first difference spectrum fitting is still the preferred method for trace
Ca quantitation. Our study also demonstrates the clear advantage of using a 200 keV system, instead of 120 keV
or lower accelerating voltages, for EELS analysis of relatively thick biological cryosections.
Key words: electron energy loss spectroscopy (EELS), Ca quantitation, internal reflection, precise correlation,
radiation damage–induced spectral changes
I NTRODUCTION
The large collection efficiency of electron spectrometers for
inelastically scattered electrons recommends electron en-
ergy-loss spectroscopy (EELS) as a valuable method for
compositional imaging with high detection sensitivity and
at high spatial resolution (Isaacson and Johnson, 1975).
Given the importance of calcium in biological signal trans-
duction, its detection and imaging has become a high-
priority objective for analytical electron microscopy in bi-
ology and it has been verified experimentally that EELS can
detect low concentrations of Ca with a 3- to 5-fold higher
sensitivity than that attainable with available X-ray detec-
tors used for electron probe microanalysis (Shuman and
Somlyo, 1987; Leapman et al., 1993; Leapman and New-
bury, 1993). Detection of Ca by EELS has been optimized
through the use of parallel detectors and implementation of
multiple least-squares fitting to extract the characteristic Ca
L-edge signal from the background (Shuman and Somlyo,
1987; Leapman and Swyt, 1988; Leapman and Andrews,
1991; Leapman and Hunt, 1991; Wang et al., 1992; Leap-
man et al., 1993; Leapman and Newbury, 1993; Door and
Ga¨ngler, 1995; Egerton, 1996), while computer modeling
Received December 17, 1998; accepted January 19, 1999.
*Corresponding author
Microsc. Microanal. 5, 17–28, 1999
MicroscopyAND
Microanalysis
© MICROSCOPY SOCIETY OF AMERICA 1999