*
E-mail: aminmokhlesi@gmail.com
Sexual Differences in Foraging Behavior of Fiddler
Crab, Uca sindensis (Decapoda: Ocypodidae)
Mokhlesi, Amin
1*
; Kamrani, Ehsan
2
; Backwell, Patricia
3
, Sajjadi, Mirmasoud
2
1- Tehran Central Branch, Islamic Azad University, Young Researchers Club, Tehran, IR Iran
2- Department of Marine Biology, Faculty of Science, University of Hormozgan,
Bandar Abbas, IR Iran.
3- School of Botany and Zoology, Australian National University, Canberra, Australia.
Received: October 2011 Accepted: December 2011
© 2011 Journal of the Persian Gulf. All rights reserved.
Abstract
Fiddler crab females have two small feeding claws while males have only one; the other is enlarged and
used as a weapon as well as a mate attraction signal. The study was conducted on the small intertidal
fiddler crab Uca sindensis in the Abi estuary near Bandar Abbas on the Persian Gulf. Data were collected
from September 2007 to August 2008 in order to cover a wide range of ecological conditions experienced
by these crabs over the duration of a year. We considered whether males Uca sindensis have behaviorally
or morphologically compensated for the loss of one feeding claw. Therefore we measured the sex ratio of
surface-active crabs; the size of the feeding claws the amount of food processed and the feeding effort
(male and female). We showed that males scoop up handfuls of sediment at approximately half the rate of
females, but spend more time feeding than do females (70.41%). Males are slightly larger than females,
but the relationship of feeding claw to carapace size is not linear: small females have relatively large
feeding claws for their size, while large males have relatively large feeding claws for their size. Males and
females process equivalent amounts of food on a daily basis. In order to compensate for the loss of one
feeding claw, males need to spend a greater amount of time feeding.
Keywords: Uca sindensis, Foraging behavior, Fiddler crab, Persian Gulf
1. Introduction
Sexual dimorphism is a common phenomenon in the
animal kingdom. Males may be brightly colored while
females are dull (e.g. bird, Butcher 1984; Slagsvold,
2000). Males may be smaller (e.g. spiders, Elgar, 1991)
or larger (e.g. most birds, reptiles and mammals,
Abouheif and Fairbairn, 1997; Andersson 1994;
Blanckenhorn, 2005; Dubey et al., 2009) than females
or may possess exaggerated appendages that are absent
in females (e.g. stag beetle, Armin, 2006; Brittany et
al., 2008). There are many studies examining sexual
dimorphisms in terms of sexual selection and
reproductive fitness, but far fewer have examined the
effect of sexually selected dimorphisms on the non-
reproductive aspects of animal life, for example, its
effect on foraging behavior and efficiency. An extreme
case of a sexually selected dimorphism affecting
feeding is found in the fiddler crab. These animals feed
on algae, bacteria and decaying plant and animal matter
in the mud surrounding their burrows (Caravello and
Cameron, 1987; Teal, 1958). They use their feeding
claws to scoop up a small amount of sediment and put
it into their mouth (Icely and Jones 1978; Miller, 1961;
Ono, 1965; Yamaguchi, 2000). Females have two small
Journal of the Persian Gulf
(Marine Science)/Vol. 2/No. 6/December 2011/7/37-44