Vol. 46 (1992) No. 2
Steam Prctreatment and Enzymatic Hydrolysis of Ccllulosic Substrates 149
Holzforschung
46(1992)149-154
Steam Pretreatment Conditions for Effective Enzymatic
Hydrolysis and Recovery Yields of Eucalyptus viminalis
Wood Chips
By L.P. Ramos
1
, C. Breuil
1
, D.J. Kushner
2
and J.N. Saddler
1
1
Forest Products Biotechnology, Faculty of Forestry, University of British Columbia, Vancouver,
CanadaV6TlW5
2
Department of Microbiology, Faculty of Medicine, University of Toronto, Toronto, Canada M5S 1A8
Keywords
Steam pretreatmcnt
Lignoccllulose
Enzymatic hydrolysis
Eucalyptus viminalis
Summary
Wood chips from Eucalyptus viminalis were steam treated over a ränge of tempcratures and rcsidence
times in the abscnce or presence of SO
2
. When the Substrate was steam treated without explosion, the
initial moisture content of the chips had a significant influence on the recovery yield and the degrce of
enzymatic hydrolysis. Prctreatment by steam explosion showed no Variation resulting from differences
in the initial moisture content of the chips. SO
2
catalysis was shown to be particulary beneficial for the
steam treatment of green chips. More than 95% of the original cellulose could be hydrolyscd to glucose
with more than 80% of the original pentosan recovered äs xylose in the water soluble fraction.This indi-
cated a need for chips with a high enough moisture content necessary to generate enough sulphurous acid
from SO
2
to act äs an effective catalyst.
Introduction
It is recognized that lignocellulosic Substrates require
some form of pretreatment if efficient enzymatic hy-
drolysis is to be achieved. A variety of physical and
chemical methods have been advocated, with the rela-
tive success of each method usually dependant on how
much of the starting materials is recovered after the
pretreatment step and the extent to which enzymatic
hydrolysis of cellulose is improved. It has been demon-
strated by various groups that steam pretreatment of
both wood and agricultural residues can be an effec-
tive method of fractionating lignocellulosic Substrates
into their major components äs well äs enhancing the
enzymatic hydrolysis of cellulose (Foody 1980; Sadd-
ler et al. 1982; Mammers and Menz 1984; Mackie et al.
1985;Dekkerefö/. 1987; Eklund et al 1988).
The pre-impregnation of wood chips with low levels
of SO
2
prior to steam treatment has been shown to im-
prove the enzymatic hydrolysis of the cellulosic com-
ponent of the Substrate äs well äs increasing the recov-
ery of sugars in the water soluble fraction (Mackie et
al. 1985; Clark and Mackie 1987; Schwald et al. 1989a,
b).There are several other advantages associated with
SO
2
impregnation. These include, the even and ef-
fective distribution of the catalyst throughout the
Substrate, enhanced survival and recovery of the
hemicellulose derived sugars and reduced tempera-
ture and residence time requirements for Optimum
steam treatment.
Holzforschung /Vol. 46 /1992 / No, 2
© Copyright 1992 by Walter de Gruyter · Berlin · New York
Although a considerable amount of work has looked
at the influence of factors such äs chip size, concentra-
tion of catalyst, need for explosion, etc., much of this
research has utilised relatively few wood species such
äs aspen (Brownell and Saddler 1984; Brownell et al.
1986) and willow (Eklund etal. 1988). When softwood
species such äs spruce (Schwald et al. 1989b) or
radiata pine (Clark and Mackie 1987) were examined
it was apparent that different pretreatment conditions
were required and that a catalyst was essential if high
cellulose hydrolysis yields were to be obtained.
During the last ten years, Brazil has established exten-
sive plantations of fast growing Eucalyptus sp. Al-
though these trees will be primarily used for pulp and
paper production, it has been proposed that wood re-
sidues from the harvesting and processing stages
could be used äs a feedstock for producing ethanol.
There has been some past work on the steam pretreat-
ment of Eucalyptus sp., however, pretreatment condi-
tions were not optimised for maximum recovery of the
hemicellulose sugars while enzymatic hydrolysis was
generally assessed after relatively high concentrations
of enzyme were added to the pretreated Substrate
(Dekker et al. 1987). In the work described in this
paper we have looked at the influence of SO
2
impre-
gnation on the recovery yield, äs a percentage of the
original Substrate, and the enzymatic hydrolysis of the
cellulosic fraction from wood chips with varying mois-
ture contents. We have identified how parameters
such äs the temperature, residence time and initial
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