292 Accepted by J. Klimaszewski: 10 Jan. 2018; published: 21 Feb. 2018 ZOOTAXA ISSN 1175-5326 (print edition) ISSN 1175-5334 (online edition) Copyright © 2018 Magnolia Press Zootaxa 4382 (2): 292–298 http://www.mapress.com/j/zt/ Article https://doi.org/10.11646/zootaxa.4382.2.4 http://zoobank.org/urn:lsid:zoobank.org:pub:AA68FBAB-EDE3-4204-A712-B7ADD0E671A9 First records of Archiconnus and Obesoconnus in Costa Rica (Coleoptera, Staphylinidae, Scydmaeninae) PAWEŁ JAŁOSZYŃSKI Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50-335 Wrocław, Poland. E-mail: scydmaenus@yahoo.com Abstract The occurrence of the glandulariinae genera Obesoconnus and Archiconnus in Costa Rica is reported for the first time. Obesoconnus was previously known from Mexico and French Guyana; Obesoconnus costaricanus sp. n. fills the gap in the hitherto known distribution of this remarkable genus. Archiconnus was known only from the northern part of Peru (east of the Andes); Archiconnus limonensis sp. n. broadens the known genus range ~ 2000 km to north-west. Key words: Glandulariini, new species, Neotropical Introduction The Scydmaeninae genus Obesoconnus Jałoszyński, 2014 comprises two species that share a very unusual body shape among Glandulariini. Typically, beetles of this tribe are approximately ant-like, i.e., they are elongate, often slender, with deep constrictions between the head and pronotum and between the pronotum and elytra. Adults of Obesoconnus are so stout that their general appearance closely resembles that of some Eutheiini, e.g., small and strongly convex species of Paraneseuthia Franz, 1986. The head capsule of Obesoconnus is unique among Glandulariini and also resembles that of Eutheiini, as it lacks the occipital constriction. However, this seems to be a secondary modification caused by enormously developed, strikingly huge composite eyes, which occupy the entire length of the exposed part of the head capsule, from mandibular bases to the 'neck' region. Other diagnostic characters include the antennae gradually thickened distally; the submentum demarcated laterally by sutures; the hypomeral ridges long but incomplete, posteriorly not reaching the gular plate; the prothorax with lateral edges; the pronotum with a transverse antebasal groove connecting 2–3 small and diffuse pits (lateral pair and a median pit; the latter may be absent) and with one pair of lateral antebasal impressions; the basisternal part of prosternum much shorter than the coxal part; procoxal cavities narrowly open; the prosternal process barely marked, with a rounded apex; the mesoventral intercoxal process carinate; the metaventral intercoxal process with a deep median notch; each elytron with one asetose rudiment of basal fovea; the aedeagus with the basal 'pumping apparatus' composed of a lentiform sclerotization located at middle of a basal membranous diaphragm and connected to an internal longitudinal apophysis providing an attachment site for muscles; and parameres present, slender. This set of characters (illustrated by Jałoszyński (2014)) makes Obesoconnus one of the most readily identifiable taxa among Glandulariini; it belongs to a large group of genera with lateral sutures demarcating the submentum, which may represent a natural group (e.g., Jałoszyński 2012a, 2013a–c, 2014, 2015a–c, 2017; Jałoszyński & Meybohm 2017). The two described Obesoconnus species inhabit French Guyana and Mexico. Archiconnus Franz, 1980 currently includes a single species from Peru, and was originally described as a taxon highly similar to Euconnus Thomson, 1859, especially to the subgenus Napochus Thomson, 1859 (Franz 1980). The label data of the holotype of A. huallaganus Franz, 1980, the type species of Archiconnus, suggest that Franz initially intended to place it within Euconnus. Later, Franz (1994) added one Mexican species to this genus. Jałoszyński (2013c) redescribed Archiconnus and maintained it as a separate genus, but he transferred the Mexican species to a new genus, Mexiconnus Jałoszyński, 2013c. The latter author provided an emended diagnosis of Archiconnus that included, among other characters, a short head with the vertex not bulging posterodorsally; thick