1022-7954/03/3903- $25.00 © 2003 MAIK “Nauka /Interperiodica” 0356
Russian Journal of Genetics, Vol. 39, No. 3, 2003, pp. 356–358. Translated from Genetika, Vol. 39, No. 3, 2003, pp. 439–441.
Original Russian Text Copyright © 2003 by Bychek, Müller.
To date, the problem of the invasion of alien or
exotic species to the territories of different countries
attracts growing attention. The importance of this prob-
lem needs no special comments. The realization of the
significance of this phenomenon resulted in the forma-
tion of working groups focused on the studies of the
estuaries and marine ecosystems of the Baltic region
and Caspian basin. Also, the international cooperation
in the field of management of the ballast waters, etc., is
being developed. The urgency of this problem for Rus-
sia can be judged from the recently organized Ameri-
can–Russian symposium on invasive or invading spe-
cies (Borok, Russia, August 27–31, 2001).
In this respect, not only studies of the biology of
newly invaded species and the ecological consequences
of their invasion, but also elucidation of their natural
distribution ranges and their migration routes become
urgent.
It should be mentioned that for a number of aquatic
invertebrate groups the problem of species identifica-
tion still remains a challenge. This primarily concerns
morphological diagnostics of some Daphnia species.
Polymorphism of the members of Daphnia longispina
complex is well known. According to Korovchinskii
[1], classification of the forms of plastic species such as
Daphnia longispina, D. galeata, and D. hyalina is
rather questionable, and there are no unambiguous cri-
teria for their isolation. This opinion is shared by
Glagolev [2], who attributes the above species to the
longispina group and considers the species isolation
within this group as provisional. Nevertheless, analysis
of Daphnia individuals from different isolated pools
resulted in establishing some reasonable morphological
differences between the above species [3]. This, how-
ever, does not solve the problem on a wider scale.
The published list of aquatic organisms inhabit-
ing the Volga River [4] contains eight Daphnia spe-
cies. D. galeata is not mentioned, although a number of
generally accepted diagnostic features indicate its pres-
ence. For these reasons, the present study was focused
on the genetic diagnostics of some frequent Daphnia
species from the Volga River.
Parthenogenetic clones were bred in laboratory con-
ditions from the round-headed (A) and helmeted (B)
Daphnia females collected in the near Damp pool of the
Kuibyshev reservoir and in the upper part of the Saratov
reservoir on May 21 and June 18, 1998.* Analogous
clones were obtained from the D. cucullata G.O. Sars
(C) individuals and from a round-headed female (D),
respectively caught on June 18 and August 30, 1998, in
the Kuibyshev reservoir. Animals were fixed in ethanol,
and total DNA was extracted from each clone by incu-
bating in the H3 buffer (10 mM Tris–HCl, pH 8.3
at 25°C; 0.05 M KCl; 0.005% NP-40; Replithern Reac-
tion Buffer, Biozym), containing 15 μg of proteinase K.
The volume of the H3 buffer depended on the number
of the animals in the sample [5].
The DNA extracts obtained were used for cloning
performed by use of polymerase chain reaction (PCR).
PCR fragments were amplified using the mitochondrial
DNA (mtDNA)(16S rRNA) specific primers, S1 (5'-
CGGCCGCCTGTTTATCAAAAACAT-3') and S2 (5'-
GGAGCTCCGGTTTGAACTCAGAT-3'), which were
the modified universal primers, LR-J-12887 and LR-N-
13398 [6]. The reaction conditions were 3 min at 93°C
for denaturation followed by 35 cycles of 92°C for 45 s;
55°C for 45 s; 72°C for 44 s; and a final incubation of
7 min at 72°C.
Molecular Genetic Diagnostics of Some Daphnia Species
(Crustacea, Cladocera) from the Volga River
E. A. Bychek
1
and J. Müller
2
1
Institute of Ecology of the Volga Basin, Russian Academy of Sciences, Togliatti, 445003 Russia;
fax: (8482)489504; e-mail: bychek@tlt.volga.ru
2
Department of Population Biology, Zoological Institute University of Mainz, Mainz, 55099 Germany;
fax: +49+6131 3923731; e-mail: jakob@oekologie.biologie.uni-mainz.de
Received March 1, 2002; in final form, August 13, 2002
Abstract—Population genetic structure of the widespread Daphnia species from Kuibyshev and Saratov res-
ervoirs was examined using RAPD–PCR technique with four mitochondrial DNA (16S rRNA) specific primers.
One of the Daphnia clones examined from the Volga region appeared to be conspecific to the North American
Daphnia galeata, while another clone was most likely a hybrid between D. galeata and D. cucullata.
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* According to the diagnostic morphological features suggested by
Manuilova [7], these Daphnia females from the Volga River are
attributed to D. longispina O.F. Müller.