REVIEW
OF
PALAEOBOTANY
AND
PALYNOLOGY
ELS EV I ER Reviewof Palaeobotanyand Palynology 84 (1995) 389-412
Cycadales and Bennettitales leaf compressions of the Bohemian
Cenomanian, Central Europe
Jifi Kva~ek
National Museum, Prague, Department of Palaeontology, V6clavskk n6m. 68, Prague 1, Czech Republic
Received 14 February 1994; revised and accepted 5 May 1994
Abstract
Three species of cycad leaves from the Bohemian Cenomanian, Central Europe, are described and revised on the
basis of their cuticle: Jirusia jirusii (Bayer) J. Kva6ek, comb. nov., Nilsonia bohemica Velenovsk~, and Nilsonia cf.
orientalis Heer, Additionally, the genus Jirusia Bayer, 1914 is newly reintroduced and defined. For the first time a
bennettitalean leaf compression species is recorded in the Bohemian Cenomanian and described as Nilssoniopteris
pecinovensis J. Kva~ek, sp. nov.
1. Introduction
The main object of this paper is to describe and
contribute to the knowledge of some new and
previously known taxa of Cycadales and
Bennettitales from the Cenomanian of Bohemia.
Fossil cycads from the Cretaceous of Bohemia
were first recorded by Reuss (1844) and Corda
(see Reuss, 1846). Many fossil cycads were
described by Velenovsk~ (1885) and Bayer (1901,
1914). While Velenovsk), who comprehensively
described the whole Cenomanian flora of Bohemia,
concentrated only on macroscopical descriptions,
Bayer (1914, 1921) used microscopical methods
and cuticle maceration. He especially collected
cuticle-bearing specimens. Further studies were
done by Velenovsk2) and Viniklfi~ (1926, 1927,
1929, 1931) and Vinikl~i~ (1933). Later, cycads
were mentioned only in lists of occurring taxa
(Knobloch, 1969, 1971, 1987).
The only previously described bennettitalean
representative from the Bohemian Cretaceous
Basin is a silicified stem, figured by N6mejc (1968).
Additionally, some leaf impressions and silicified
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stems are known from the Lower Cretaceous
of Moravia (Hlugtik and Purkyfiovfi, 1982;
Purkyfiovfi, 1983).
2. Material and methods
Material for the present study has been
derived from the Peruc-Korycany Formation
(Cenomanian) of the Bohemian Cretaceous Basin,
as defined by Cech et al. (1980). The Peruc-
Korycany Formation consists of the Peruc
Member, the Korycany Member and the Pecinov
Member (Fig. 1). The lower part of the Peruc
Member includes fluvial sands and sandstones,
mudstones and interbedded mudstones, supratidal
marsh and back swamp deposits crossed by mean-
dering tidal creeks close to the eustuarine mouth
in the area of south-central Bohemia (Uli6n~,
1992). It continually evolved into the marine
Korycany Member, built by sandstones containing
a rich marine fauna and occasionally preserved
stems of tree ferns and poor leaf impressions. In