probably is of E. fragilis. If it were designated as the lectotype of
Host’s name, E. major would have been widely and persistently used
in the sense of E. nebrodensis, i.e., not including the type of E. major,
and then Art. 57.1 of the ICN would need to be applied.
We might propose the name E. major for conservation under Art.
14.1 of the ICN with a conserved type belonging to E. nebrodensis.
However, this would not clarify the confused nomenclatural history
of the name E. major. Therefore, we propose instead its rejection under
Art. 56.1.
Author information
SB, https://orcid.org/0000-0003-2568-7278
EDG, https://orcid.org/0000-0001-9349-1328
Acknowledgements
We are grateful to N. Turland (Botanic Garden and Botanic
Museum, Freie Universität, Berlin) and John McNeill (Royal Botanic
Garden, Edinburgh, Scotland, U.K. & Royal Ontario Museum,
Toronto, Canada) for useful suggestions and editorial comments.
(2805) Proposal to conserve Eulophia, nom. cons., against the additional name
Geodorum (Orchidaceae: Eulophiinae)
Mark W. Chase,
1,2
Maarten J.M. Christenhusz,
2
Pankaj Kumar
3
& André Schuiteman
1
1 Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, U.K.
2 Department of Environment and Agriculture, Curtin University, Perth, Western Australia, Australia
3 Kadoorie Farm and Botanic Garden, Lam Kam Road, Tai Po, New Territories, Hong Kong S.A.R., P.R. China
Address for correspondence: Mark W. Chase, m.chase@kew.org
DOI https://doi.org/10.1002/tax.12480
First published as part of this issue. See online for details.
(2805) Eulophia R. Br. in Bot. Reg.: ad t. 573 (‘578’). 1 Nov 1821
(‘Eulophus’)[Orchid.], nom. et orth. cons.
Typus: E. guineensis Lindl. (in Bot. Reg.: t. 686. 1 Feb
1823), typ. cons.
(=) Graphorkis Thouars in Nouv. Bull. Sci. Soc. Philom. Paris 1:
318. Apr 1809 (nom. cons.) (etiam vide), nom. rej.
(=) Geodorum Andrews in Bot. Repos.: ad t. 626. Jun 1811, nom.
rej. prop.
Typus: G. citrinum Andrews
(=) Lissochilus R. Br. in Bot. Reg.: ad t. 573 (‘578’). 1 Nov 1821,
nom. rej.
Typus: L. speciosus R. Br.
Phylogenetic analyses of the orchid subtribe Eulophiinae
(Martos & al. in Taxon 63: 9–23. 2014; Bone & al. in Bot. J. Linn.
Soc. 179: 43–56. 2015) have clearly demonstrated that the genus
Eulophia as currently circumscribed is not monophyletic, with
Acrolophia Pfitzer (Entwurf Anordn. Orch.: 59. 1887), Geodorum
Andrews (in Bot. Repos.: ad t. 626. 1811), Oeceoclades Lindl.
(in Edwards’s Bot. Reg.: ad t. 1522. 1832) and several other genera
embedded. Most names in the other embedded genera exist in Eulo-
phia, but not for species published as Geodorum, an older name.
Eulophia in the broad sense (including Acrolophia and Oeceoclades)
is a relatively large genus (c. 250 species; Chase & al. in Bot. J. Linn.
Soc. 177: 151–174. 2015). It has a pantropical distribution, but with
the greatest species diversity in Africa where it is important in many
floras. It figures to a minor extent in horticulture. Geodorum is found
in tropical Asia and Australasia and has only 9 species (Chase & al.,
l.c.), none commonly found in horticulture. It differs from Eulophia
in the entire lip (versus mostly lip 3-lobed) and in its inflorescence
structure: the flower-bearing apex is nodding. To avoid the need to
transfer Oeceoclades and Acrolophia to Eulophia, Martos & al.
(l.c.) and Bone & al. (l.c.) proposed to segregate Orthochilus Hochst.
ex A. Rich. (Tent. Fl. Abyss. 2: 284. 1850) from Eulophia, emphasiz-
ing that the sepals of these species are similar in colour to the petals,
whereas those in Eulophia s.str. are dissimilar. However, this differ-
ence and others mentioned by those authors also occur often in Eulo-
phia s.str., if comparisons are made with the Asian/Australasian
species. Even if exclusion of the genus Orthochilus as advanced by
Martos & al. (l.c.) and Bone & al. (l.c.) is accepted, the issue of the
priority of Geodorum over Eulophia s.str. would still need to be
addressed because Bone & al. (l.c.) found Geodorum to be deeply
embedded in a clade that includes the type of Eulophia. Indeed these
authors (l.c.: 53) indicated their submission of a proposal to conserve
the latter name over the former; however, no such proposalwas ever
submitted.
We advocate instead a broad generic concept of Eulophia
including all these genera, all of which, except Geodorum, have been
included by at least some authors in Eulophia in the past. Geodorum
preceded Eulophia by 10 years, and both names are still in common
© 2021 The Authors.
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432 Version of Record
Chase & al. • (2805) Conserve Eulophia TAXON 70 (2) • April 2021: 432–433