i
i
Forest
Forest
Biogeosciences and Forestry Biogeosciences and Forestry
Ectomycorrhizal fungal community in mature white poplar plantation
Marina Milović
(1)
,
Saša Orlović
(1)
,
Tine Grebenc
(2)
,
Marko Bajc
(2)
,
Branislav Kovačević
(1)
,
Hojka Kraigher
(2)
Ectomycorrhizal communities are rarely studied on seasonal basis, especially
in poplar plantations. In this study we analysed the ectomycorrhizal commu-
nity in a mature twenty-year-old white poplar ( Populus alba L.) plantation
during four consecutive seasons. Using morpho-anatomical and molecular
identification 30 taxa of ectomycorrhizal fungi were recorded of which 15
were identified to the species level, 12 to the genus level, 2 to the family, and
one morphotype of ectomycorrhizae remained unidentified. The most abun-
dant among identified ectomycorrhizal fungi were: Inocybe griseovelata, Ino-
cybe splendens, Tuber rufum, and Tomentella sp. 2, which together repre-
sented up to 50% of all ectomycorrhizal root tips. The number of ectomycor-
rhizal fungal taxa and the percentage of vital ectomycorrhizal root tips were
highest in winter and spring, respectively. The diversity indices of ectomycor-
rhizae, number of vital ectomycorrhizal root tips, and total fine roots in the
studied poplar plantation did not differ between seasons. Ectomycorrhizal
fungi belonging to Inocybaceae family and the short-distance exploration strat-
egy were dominant in all four seasons. On the other hand, the abundance of
ectomycorrhizal root tips belonging to the medium-distance exploration strat-
egy type was significantly higher in spring in comparison with autumn and win-
ter.
Keywords: Populus alba L., Ectomycorrhizal Diversity, Morpho-anatomical
Characterization, Molecular Identification, Seasons
Introduction
White poplar (Populus alba L.) is a pioneer
tree species frequently found in lowlands,
in humid areas, along rivers, and in flood
land forests. The species has a wide distri-
bution range from the Mediterranean to
central Asia (Palancean et al. 2018). It has
relevant environmental functions, such as
prevention of soil erosion and formation of
windbreaks, as well as ecological impor-
tance for maintaining the high biodiversity
of riparian woodland. Poplars are fast-
growing species with strong regenerative
abilities, whose wood is used for the pro-
duction of pellets and energy as well as
raw material for the pulp and paper indus-
try (Palancean et al. 2018). Poplars regu-
larly form dual associations with ectomyc-
orrhizal (ECM) fungi and arbuscular mycor-
rhizal fungi which are known to prefer dif-
ferent soil conditions (Karlinski et al. 2010,
Katanić et al. 2013). A detailed review on
ectomycorrhiza of Populus was given by
Szuba (2015).
Mycorrhizal fungi essentially contribute
to the functioning of the forest ecosystem,
biogeochemical processes, and forest pro-
ductivity. They obtain water and nutrients
from the soil and deliver them to the host
plant in exchange for photosynthetically-
produced carbohydrates (Smith & Read
2008). The ECM fungi are able to access
limiting nutrients (usually nitrogen) situ-
ated beyond the nutrient depletion zone
which surrounds the root system of the
host plant (Smith & Read 2008). Also, they
can release nitrogen from immobilized
sources normally inaccessible to the plant,
such as soil organic matter or leaf litter
that are accumulated on the forest floor.
Additionally, they have an important role in
promoting carbon sequestration in soil
through increasing carbon storage (Stuart
& Plett 2020). Mycorrhizal fungi link above-
ground and belowground components of
forest ecosystems through the common
mycelial network which increases seed-
lings’ survival and improves their physiol-
ogy, as seedlings can receive carbon, nutri-
ents, and water from the older trees (Sel-
osse et al. 2006). They can improve plant
tolerance to different abiotic stress factors
as well as boost plant immunity and in-
crease resistance to pathogens (Smith &
Read 2008, Milović et al. 2021). Further-
more, mycorrhizas have an important role
in forest ecosystems’ stability under cli-
mate change, by mitigating the detrimental
effects of different unfavorable factors as
increased CO
2
in the atmosphere, tempera-
ture rise, drought, soil acidification, pollu-
tants, pests, diseases, etc. (Milović et al.
2021).
Since the functional compatibility and
stress tolerance of ectomycorrhizae de-
pend on both partners, especially on fun-
gus, the information on the ECM commu-
nity structure can be valuable in studies of
forest ecosystems functioning (Kraigher
1996).
ECM fungal communities on autochtho-
nous white poplars were investigated in a
drought-adapted natural forest in the Hun-
garian Plain in Hungary (Jakucs 2002), and
in a natural poplar stand in Special Nature
Reserve “Kovilj-Petrovaradin marshes” in
Serbia (Katanić et al. 2015b). Worldwide
ECM fungal communities associated with
Populus spp. in nature forest were studied
© SISEF https://iforest.sisef.org/ 540 iForest 14: 540-547
(1) University of Novi Sad, Institute of Lowland Forestry and Environment, Antona Čehova
13, 21000 Novi Sad (Serbia); (2) Slovenian Forestry Institute, Večna pot 2, 1000 Ljubljana
(Slovenia)
@ Marina Milović (katanicm@uns.ac.rs)
Received: Mar 26, 2021 - Accepted: Sep 24, 2021
Citation: Milović M, Orlović S, Grebenc T, Bajc M, Kovačević B, Kraigher H (2021).
Ectomycorrhizal fungal community in mature white poplar plantation. iForest 14: 540-547. –
doi: 10.3832/ifor3827-014 [online 2021-11-26]
Communicated by: Maurizio Ventura
Research Article Research Article
doi: doi: 10.3832/ifor3827-014 10.3832/ifor3827-014
vol. 14, pp. 540-547 vol. 14, pp. 540-547