Review Article
Volume 3 Issue 4- September 2018
Ann Rev Resear
Copyright © All rights are reserved by Fábio Pereira Leivas Leite
Equine Theileriosis: Review
Ana Muñoz Vianna
1,2
, Ana Paula de Souza Stori de Lara
3
, Guilherme Borges Weege
3
, Rodrigo Casquero Cunha
2
and Fábio Pereira Leivas Leite
1,2
*
1
Department of Microbiology and Parasitology, Federal University of Pelotas (UFPel), Brazil
2
Postgraduate Program in Veterinary Medicine, Federal University of Pelotas (UFPel), Brazil
3
Department of Microbiology and Parasitology, Federal University of Pelotas (UFPel), Brazil
Submission: June 06, 2018; Published: September 28, 2018
*
Corresponding author: Fábio Pereira Leivas Leite, Veterinary Graduate Program Federal University of Pelotas - UFPel, Capão do Leão Campus,
Pelotas, RS, Brazil, Tel: ; Email:
Introduction
Equine piroplasmosis, caused by the intracellular protozoa
Theileria equi and Babesia caballi, belonging to the phylum
Apicomplexa, is considered the most important horse disease
transmitted by ticks [1]. The disease is caused by infection with
Theileria equi, formerly Babesia equi [2] and is called equine
theileriosis [3,4]. The phylum Apicomplexa includes a large group
of organisms, including the order Piroplasmida, comprised
of compulsory vertebrate and invertebrate parasites [5]. The
organisms have an apical complex containing secretory organelles
involved in invasion and the parasite remains within the host
[5]. Theileria equi has a wide geographic distribution and infects
horses in Europe, Asia, Africa and South and Central America,
but is considered endemic in tropical and subtropical [6]. Equine
theileriosis stands out as a major cause of both horse illness and
of losses to the horse raising industry. Even with improvements
in health, piroplasmosis causes losses associated with clinical
factors and through international restrictions on the transit of
seropositive animals [7,8]. Theileria equi is small, 2-3 mm in
length [9], pleomorphic within erythrocytes, and is isolated in
pairs or when forming tetrads (Maltese cross). When parasitemia
is low, they are barely visible under the microscope and can be
easily confused with technical artifacts due to their small size [7].
The parasite is carried by ticks, especially the genera Anocentor,
Amblyomma, Rhipicephalus and Haemaphysalis [10]. Rhipicephalus
microplus is considered the responsible for infestations in horses
when sharing pastures with bovine [11,12].
When taking a blood meal on a host, (an infected equine),
the tick vector ingests erythrocytes containing the gametocytes
that in its intestines will give rise to the gametes that will fuse
to form the zygote. The zygote differentiates to the mobile form
ookinete which is transported through the hemolymph to the
salivary glands of the tick. Within the salivary glands, sporoblasts,
and finally, sporozoites will give rise (MEHLHORN & SCHEIN,
1998.Theileria equi transmission occurs through tick blood meals
which contain infective (sporozoite) forms (UILENBERG, 2006).
Theileria equi exhibits a biphasic cycle in the mammalian host
with an intra-leucocyte development phase followed by an intra-
erythrocytes phase (RAMSAY et al., 2013). The intra-leucocyte
phase includes infection of B and T lymphocytes, monocytes,
and macrophages. However, unlike others theilerias, the pre-
erythrocytic stage of T. equi has not been clinically associated
with disease, since it is not necessary to complete the lifecycle
Ramsay et al. 2013. Approximately 2 days after infection T. equi
schizonts begin to infect lymphocytes, and by 15 days merozoites
can be viewed through blood smears in erythrocytes. The
parasitemia reaches approximately 7% of the erythrocytes, but in
immunocompromised animals it can reach 80% [14]. In nature,
horses are T. equi reservoirs where transstadial transmission
can occur [15]. In transstadial transmission, larvae that feed on
infected horses are infected yet maintain sporozoites in their
salivary glands. When changing into nymphs, these will then
Ann Rev Resear 3(4): ARR.MS.ID.555620 (2018) 00100
Annals of
Reviews and Research
Abstract
Equine theileriosis, caused by the Apicomplexa protozoan Theileria equi, is an intra-erythrocytic parasite disease found in horses, and it is
widely distributed throughout the world. Its prevalence and endemicity are linked to the presence of tick vectors and chronic equine carriers.
Diagnosis of theileriosis is based on serological methods, including ELISA (Enzyme-Linked Immunosorbent Assay), IFAT (indirect fluorescent
antibody), and molecular methods such as PCR (Polymerase Chain Reaction). Theileria equi has merozoite surface proteins, equi merozoite
antigens (EMA-1, EMA-2, EMA-3 and EMA-6). These antigens play an important role in pathogenesis and provide as host immune response
targets against the parasite. EMA-1 and EMA-2 have been identified as immunodominant antigens and they are expressed on the surface of
extra-erythrocyte merozoites. EMA-1 protein has been used in commercial testing of immunodiagnostic; however, EMA-2 is a highly conserved
immunogenic protein and a promising target. The aim of this study is to contribute to improving the diagnosis and prevention of equine
theileriosis.
Keywords: ELISA; Immunodiagnostics; Surface protein and Theileria equi