© Senckenberg Museum of Natural History Görlitz · 2015
ISSN 1864-6417
pp. 85–89
SHORT COMMUNICATION
Short note on enchytraeid occurrence in deep layers of urban soils
Jiří Schlaghamerský
Masaryk University, Faculty of Science, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic
E-mail: jiris@sci.muni.cz
Received 27 February 2015 | Accepted 10 June 2015
Published online at www.soil-organisms.de 1 August 2015 | Printed version 15 August 2015
87 (2) · August 2015
1. Introduction
Most soil organisms require aerobic conditions and
access to carbon as a source of energy and building
material for their bodies. Therefore, in soils of most
terrestrial ecosystems animals are concentrated in
the upper 10–15 cm layer and few individuals of few
species occur below 50 cm depth. Where plant roots or
earthworm tunnels reach further down, also other soil
organisms follow along these rather thin structures, i.
e. the rhizo- and drilosphere (Lavelle & Spain 2005).
Exceptions where a more general colonization of soil into
greater depth occurs are dry grasslands and arable fields.
Ploughing relocates substantial amounts of dead organic
matter into greater depth whereas the upper soil is more
exposed to desiccation and freezing, resulting in a more
even distribution of soil organisms along the vertical
profile and their penetration in considerable numbers at
least down to the plough pan. Also buried soil horizons
with adequate dead organic matter content might host
soil organisms as long as some exchange of soil air
provides enough oxygen (Gurwick et al. 2008). Such
buried horizons are typical of alluvial soils. Buried soil
horizons or buried layers or pockets of humus-rich soil
or of other organic materials can also be found in urban
soils as a consequence of land reclamation, landscaping,
construction activities, dumping of waste etc. However,
most samplings for soil fauna have been limited to the
upper soil layer of some 8 to 15 cm based on the general
experience that further down numbers of soil fauna are
negligible (and also due to the fact that extracting soil
cores reaching further down can get difficult even in
deeply weathered soils). This all applies also to the vertical
distribution of potworms or enchytraeids (Annelida:
Clitellata: Enchytraeidae), that are a numerically and
functionally important part of soil mesofauna. Very few
data are available about the occurrence of enchytraeids
below 20 cm soil depth (see Discussion). Although
enchytraeids reach only low densities in deep soil and
their occurrence is clustered based on the availability
of sources of organic matter (Dózsa-Farkas 1992), their
occurrence still shows that they find food and participate
in decomposition processes at substantial depth. Deeper
layers might also host specialized species hitherto
unknown to science (Dózsa-Farkas 1991).
As part of an environmental assessment project,
samples of anthroposols were taken from several sites in
the Swedish city of Malmö at three different depths down
to 2 m below ground. Soil samples were examined to find
out if they contained enchytraeids and if so what species
and in what numbers were present. Unfortunately, only
little information about the project and sampling sites
was available to the author; presented background data
are mostly based on Törneman (2014). Nevertheless this
was a rare opportunity to investigate the possibility of
soil animal live in urban soils at substantial depth.
2. Material and methods
In October 2012, five sites in the city of Malmö in
southern Sweden were sampled, covering reclaimed
land areas at the sea front as well as old industrial and
residential land, currently used as urban greens of
various types and sizes (Tab. 1). At each site one pit was
dug and soil was sampled from layers defined by three
depth ranges: 0.1–0.5 m, 0.6–1.0 m and 1.6–2.0 m (for the
exact ranges actually sampled at the individual sites see
Tab. 1). The individual samples were put into plastic