International Journal of Neuroscience, Early Online, 1–6, 2012
Copyright © 2012 Informa Healthcare USA, Inc.
ISSN: 0020-7454 print / 1543-5245 online
DOI: 10.3109/00207454.2012.678444
miR-15a and 16-1 Are Downregulated in CD4
+
T Cells of
Multiple Sclerosis Relapsing Patients
Julio Cesar Cetrulo Lorenzi,
1,2
Doralina G. Brum,
3
Dalila L. Zanette,
1,2
Alessandra de Paula
Alves Souza,
2,4
Fernanda Gonc ¸alves Barbuzano,
2
Antonio Carlos dos Santos,
5
Amilton Antunes Barreira,
3
and Wilson Araujo Silva, Jr.
1,2
1
Genetics Department, S˜ ao Paulo University Medical school of Ribeir˜ ao Preto, Ribeir˜ ao Preto, Brazil
2
National Institute of Science and Technology in Stem Cell and Cell Therapy (INCTC), Ribeir˜ ao Preto, Brazil
3
Neurosciences Department, S˜ ao Paulo University Medical School of Ribeir˜ ao Preto, Ribeir˜ ao Preto, Brazil
4
Immunology Department, S˜ ao Paulo University Medical School of Ribeir˜ ao Preto, Ribeir˜ ao Preto, Brazil
5
Internal Medicine Department, S˜ ao Paulo University Medical School of Ribeir˜ ao Preto, Ribeir˜ ao Preto, Brazil
ABSTRACT
The pathology of relapsing–remitting multiple sclerosis (RR-MS) is largely attributed to activated autoreactive
effector T lymphocytes. The inluence of microRNAs on the immune response has been shown to occur in
different pathways of lymphocyte differentiation and function. Here, the expression of the miRNAs miR-15a/16-
1 in PBMC, CD4
+
, and CD8
+
from RR-MS patients has been investigated. BCL2, a known miR-15a/16-1 target,
has also been analyzed. The results have shown that miR-15a/16-1 is downregulated in CD4
+
T cells, whereas
BCL2 is highly expressed in RR-MS patients only. Our data suggest that miR-15a/16-1 can also modulate the
BCL2 gene expression in CD4
+
T cells from RR-MS patients, thereby affecting apoptosis processes.
KEYWORDS: apoptosis, BCL-2, lymphocytes, miRNAs
INTRODUCTION
Multiple sclerosis (MS) is a demyelinating inlamma-
tory disease of the central nervous system (CNS). The
relapsing–remitting (RR) course is the most frequent
clinical presentation of MS. Relapses are related to in-
lammatory lesions that can be detected by magnetic
resonance imaging (MRI) [1]. The pathology of MS is
largely attributed to activated autoreactive T-cell effec-
tors, mainly CD4
+
, that migrate to the CNS through
the blood–brain barrier. The inlammation caused by
T lymphocytes is especially persistent in MS. This can
be due to the increased proliferative capacity of autore-
active cells [2]; the enhanced expression of apoptosis
Received 9 November 2011.
These authors contributed equally to this work.
We thank Anemari Ramos Dinarte dos Santos for the technical support. This
work has been supported by FAPESP (2010/00310-1, 1998/14247-6), CNPQ,
CAPES, and FUNDHERP.
Address correspondence to Wilson Araujo da Silva Jr., Centro Regional de
Hemoterapia de Ribeir˜ ao Preto, Rua Cat˜ ao Roxo, 2501, Monte Alegre, CEP
14051-140, Ribeir˜ ao Preto, Brasil. E-mail: wilsonjr@usp.br
inhibitor (IAP) family members IAP, IAP-2, and X-
IAP [3]; and the higher expression of anti-apoptotic
molecules, such as BCL-2 [4, 5]. The BCL-2 gene has
important functions in cell survival and death, since it
controls the integrity of mitochondrial membranes and
the release of pro-apoptotic molecules from the mito-
chondria [6, 7]. In this way, the inlammatory T cells
present in MS become more resistant to apoptotic stim-
uli, thereby leading to persistent tissue inlammation.
The differential gene expression data regarding MS
show that immune-related genes such as CXCR5,
IL2RA, IL7R, IL7, IL12RB1, and IL22RA2 are related
to MS [8–10]. The concepts involving gene expression
regulation have evolved after the introduction of a new
class of RNA, the microRNAs (miRNAs) [11], which
are small RNAs that negatively regulate gene expression
by degradation of mRNA targets or by interruption of
translation. The inluence of miRNA on the immune re-
sponse has been demonstrated in many processes, such
as lymphocyte differentiation [12] and T regulatory cell
function [13]. The miRNA-15a and 16-1 cluster con-
trols apoptosis by targeting BCL2, as observed by the
1
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