On the Interaction of Non-Thermal Atmospheric Pressure Plasma with Tissues
S. Kalghatgi, C. Kelly, E. Cerchar, R. Sensenig, A. Brooks, A. Fridman, A. Morss-Clyne,
J. Azizkhan-Clifford, G. Friedman
Drexel University, 3141 Chestnut St, Bossone 404
Philadelphia, PA, USA
Abstract
Non-thermal atmospheric pressure plasma is
now being widely developed for various clinical
applications such as skin sterilization, blood coagulation,
cancer treatment, angiogenesis and wound healing among
others. However, understanding of mechanism of
interaction between non-thermal plasma and mammalian
cells is lacking. Here we investigated the possibility that
the dose of non-thermal plasma can be tuned to achieve
various results depending on the clinical applications
ranging from enhanced cell proliferation to inducing
apoptosis in malignant tissue. We also present some of
the underlying mechanisms of interaction of non-thermal
plasma with mammalian cells.
I. INTRODUCTION
Thermal plasma has been employed in medicine for
coagulation and ablation for some time [1]. Treatment of
tissues and cells by non-thermal plasma, where the gas
temperature is nearly at room temperature, is a recent
development [2]. It has been noted that non-thermal
plasma applied directly to surfaces of living tissues can
coagulate blood; however, it does so without charring the
tissue [2, 3]. Similarly, non-thermal plasma appears to
kill bacteria on the surface of living tissue without
histologically visible damage [2]. It has been reported
that non-thermal plasma can also mediate attachment of
cells to substrates [4-6], increase transfection efficiency
[7, 8] and surface sterilization [9-12]. Ability to tune non-
thermal plasma effects together with the simplicity of
plasma generating devices and localized nature of plasma
application makes it a promising tool in medicine.
However, mechanisms of interaction between non-
thermal plasma and living systems have been poorly
understood.
Here we study the mechanisms of interaction between
non-thermal plasma and mammalian cells. Several
different methods of non-thermal plasma generation at
atmospheric pressure are known. The type of non-thermal
plasma employed in this study is called the Dielectric
Barrier Discharge (DBD). It was invented by Siemens in
1859 [13]. The plasma in this discharge is created when
the time-varying high voltage reaches sufficient
magnitude to cause air breakdown. The presence of
dielectric layer (dielectric barrier) in the path of the
discharge limits its current which, in turn, limits the
energy transferred to ions and neutral gas species keeping
their temperature low. Although the plasma gas
temperature is low, the presence of charged particles,
radicals and electronically excited molecules and atoms
makes DBD plasma a potentially active medium whose
properties can be controlled to some extent through gas
composition as well as waveform of the time-varying
applied voltage.
II. METHODS AND MATERIALS
Mammalian Breast Epithelial Cells (MCF10A) were
maintained in high glucose Dulbecco’s Modified Eagle’s
Medium-Ham’s F12 50:50 mixture (DMEM-Ham’s F12
50:50) (Cellgro, Mediatech, VA, USA) supplemented
with 5% donor horse serum (Sigma Aldrich, St. Louis,
MO, USA), Epidermal Growth Factor (EGF, 100 µg/ml,
Sigma Aldrich, St. Louis, MO, USA), Hydrocortisone (1
mg/ml, Sigma Aldrich, St. Louis, MO, USA ), Cholera
Toxin (1 mg/ml), Insulin (10 mg/ml, and
Penicillin/Streptomycin (500 μl, 10000 U/ml penicillin
and 10 mg/ml streptomycin). Media was changed every
two days. For plasma treatment, cells were washed with
phosphate buffered saline (PBS), detached with 0.25%
trypsin (GIBCO, Invitrogen, CA, USA), and seeded near
confluence (4 x 10
5
cells/well) on 22 x 22 mm square
glass cover slips (VWR, PA, USA ) in 6-well plates
(Greiner Bio One, NC, USA). Cells were cultured for 24
hours prior to plasma treatment in 2.0 ml supplemented
media in a 37°C, 5% CO
2
incubator to allow full
attachment and spreading.
N-Acetyl-L-cysteine (2 mM, Sigma-Aldrich, St Louis,
MO), an intracellular reactive oxygen species (ROS)
scavenger and sodium pyruvate (10 mM, Sigma-Aldrich,
St Louis, MO), an extracellular ROS scavenger were used
to block the reactive oxygen species produced by non-
thermal plasma treatment.
A. Plasma Treatment
Non-thermal atmospheric pressure dielectric barrier
discharge plasma was produced using an experimental
setup similar to one previously described and
schematically illustrated in Figure 1 B. The non-thermal
plasma was generated by applying alternating polarity
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