J. Zool., Lond. (1988) 216, 519-528
Dentition and diet in snakes: adaptations to oophagy in the
Australian elapid genus Simoselaps
J. D. SCANLON
1
AND R. SHINE
Zoology A08, The University of Sydney, NSW 2006, Australia
(Accepted 23 February 1988)
(With 2 figures in the text)
Most small fossorial proteroglyphous Australian snakes of the genus Simose/aps feed on adult
lizards, but the species of one lineage (the semifasciatus group) feed exclusively on the eggs of
squamate reptiles. Examination of cleared, alizarin preparations showed that dentition of the
saurophagous species is similar to that of other elapids, but dentition of the oophagous taxa is
highly modified . The anterior (palatine and maxillary) teeth other than the fangs are reduced in
size and number whereas those of the pterygoid (and in S. 'roperi', the dentary) are enlarged
posteriorly, becoming compressed along a longitudinal plane and angled media lly . The shape of
the pterygoid and quadrate is also modified.
Two Simose/aps species with broader diets (eating both adult lizards and their eggs) show
typical 'saurophagous' dentition in one case, 'oophagous' dentition in the other, showing that
either type of dentition can be used to capture and ingest either type of prey. We suggest
functional explanations for the dentitional modifications in the egg-eating snakes, primarily in
terms of the advantages of applying considera ble force to the eggshell. Oophagous modifications
within Simose/aps are convergent with those seen in several independently-derived lineages of
oophagous colubrid snakes, but (perhaps because of the presence of the fang) differ in having the
enlarged blade-like teeth on the pterygoid or dentary rather than the maxilla.
Introduction
Materials and methods
Results
Discussion
References ..
Contents
Introduction
Page
519
520
521
525
527
Many species of snakes feed on only one or a few types of prey, and show corresponding
modifications of cranial anatomy and dentition. Distantly related taxa of snakes often show
remarkable convergences in trophic morphology (e .g. Savitsky, 1983). The best-studied example
of this phenomenon is the independent evolution of enlarged venom-conducting fangs in several
lineages of snakes (e .g. McDowell, 1986), but even more detailed resemblances have evolved
among snakes with highly specialized diets. Perhaps the best example occurs among snakes which
feed mainly or exclusively on reptilian eggs. Reduction in size and number of the anterior
I Present address: Department of Zoology, University of New South Wales, P.O. Box I, Kensington
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