Biogeography of Neurergus strauchii barani Öz, 1994 and N. s.
strauchii (Steindachner, 1887) (Amphibia: Salamandridae) assessed
using morphological and molecular data
Frank Pasmans
1
, Sergé Bogaerts
2
, Tonnie Woeltjes
3
, Salvador Carranza
4
Abstract. Five new locations of N. strauchii are reported from Turkey, closing the gap between the two subspecies N. s.
barani and N. s. strauchii. A molecular analysis based on 829 base pairs from two mitochondrial ribosomal genes (12S and
16S rRNA), together with geographical data from the area concerned, indicate all new populations found are very closely
related to the mitochondrial sequences from specimens of N. s. strauchii from its type locality and suggest the river Euphrates
might have acted as a natural barrier separating the only populations of N. s. barani known to date from all other populations
of N. s. strauchii. A morphological analysis of all N. strauchii populations sampled for this study indicates that belly patterns
are not a good diagnostic character to differentiate between N. s. barani and N. s. strauchii, suggesting the only reliable
morphological taxonomic character that allows their identification is the different number of yellow spots in adult specimens,
which is significantly higher in N. s. strauchii. The analyses also indicate that the number of spots is similar between sub-adult
and adult N. s. barani but significantly different between sub-adult and adult specimens of N. s. strauchii, suggesting there is
also a difference between both subspecies in increase in the number of spots during maturation.
Listed by IUCN as vulnerable and protected by the Bern Convention (1979), our findings indicate N. s. strauchii is more
widely distributed than previously thought, while N. s. barani would be restricted to just a few localities. Disturbance of its
prime habitat, mountain brooks, and uncontrolled illegal collection, especially of N. s. barani, are still the main threats to the
Anatolia newt throughout its known distribution range.
Introduction
The genus Neurergus is represented by two
species in Turkey: Neurergus strauchii (Stein-
dachner, 1887) and N. crocatus Cope, 1862
(Baran and Öz, 1986). The nominate sub-
species N. s. strauchii (Steindachner, 1887) is
known from the lake Van area (Schmidtler and
Schmidtler, 1970). In 1994, the subspecies N. s.
barani Öz, 1994 was described from the moun-
tains near the city of Malatya (Öz, 1994).This
subspecies is characterized by scarcity of the
dorsal yellow spots and the continuity of the
ventral orange line from the gular region to the
1 - Department of Pathology, Bacteriology and Avian Dis-
eases, Faculty of Veterinary Medicine, Ghent Univer-
sity, Salisburylaan 133, B-9820 Belgium
e-mail: frank.pasmans@UGent.be
2 - Honigbijenhof 3, NL-6533 RW Nijmegen, The Nether-
lands
3 - Molenweg 43, NL-6542 PR Nijmegen, The Netherlands
4 - Departament de Biologia Animal, Universitat de
Barcelona, Av. Diagonal 645, E-08028, Barcelona,
Spain
tip of the tail (Öz, 1994). Moreover, N. s. barani
larvae show more characteristics of pond-type
larvae compared to the larvae of N. s. strauchii
(Steinfartz, 1995). Both subspecies are sepa-
rated by a gap of approximately 300 km, from
which no specimens are known. A genetic study
using mitochondrial gene sequences, allozymes
and three plasma protein loci from all known
species of Neurergus ratified the subspecific sta-
tus of N. s. barani from the genetic point of view
and suggested this subspecies split from N. s.
strauchii approximately 3 mya (Steinfartz et al.,
2002).
The main aim of this study was to deter-
mine the possible presence of populations of N.
strauchii in the area between the two previously
known areas (populations 1, 6 and 7 for N. s.
barani and 2 and 8 for N. s. strauchii in fig. 1),
to collect data on their morphology and ecology
and to assess the level of molecular and mor-
phological differentiation among them.
© Koninklijke Brill NV, Leiden, 2006. Amphibia-Reptilia 27 (2006): 281-288
Also available online - www.brill.nl