Hydrobiologia 474: 125–130, 2002.
© 2002 Kluwer Academic Publishers. Printed in the Netherlands.
125
Field experiments on colonization and movements of stream invertebrates
in an Apennine river (Visone, NW Italy)
Stefano Fenoglio, Paolo Agosta, Tiziano Bo & Marco Cucco
University of Eastern Piedmont, Dept. Sciences and Adv. Tech., via Cavour 84, 15100 Alessandria, Italy
Tel: 131-283-835. Fax: 131-254-410. E-mail: fenoglio@unipmn.it
Received 15 June 2001; accepted 31 January 2002
Key words: macroinvertebrates, lotic environments, recolonization movements
Abstract
Macroinvertebrates continuously redistribute themselves in the riverbed. A knowledge of the colonization mech-
anisms and movement patterns is very important for an understanding of processes of restoration of lotic
environments, particularly of inland waters with severe pollution. We tested the colonization patterns of stream
macroinvertebrates in the Visone River, a tributary of the highly contaminated Bormida River (NW Italy). We
placed six groups of traps in the riverbed, each group consisting of three traps: the C trap allowed colonization
from all directions, while the D and U traps allowed access only from downstream and upstream respectively.
The C traps were the most colonized substrates, both in number of individuals and taxa. The U traps were more
colonized than the D traps, demonstrating the great importance of movements directed downstream. We report data
on taxonomic and seasonal differences in the colonization process.
Introduction
Benthic macroinvertebrates are a central element of
lotic environments, playing an important role in the
processing of allochthonous and autochthonous or-
ganic substances and in the river’s self-purifying ca-
pacity (Vannote et al., 1980). Furthermore, their
community structure is widely used as an indicator
in environmental quality monitoring (Ghetti, 1997).
It has long been known that freshwater invertebrate
populations continually redistribute themselves over
the stream bed. Environmental alterations can deplete
or destroy a benthic biocenosis, but a recolonization
process begins as soon as conditions are restored.
Many studies have shown that macroinvertebrates can
quickly colonize new or disturbed substrates from
source areas. Recolonization studies in running water
habitats have shown that animals promptly reappear in
affected areas (Williams, 1980). Colonization depends
on many elements, like invertebrate mobility, substrate
texture (Wise et al., 1979) and associated food sup-
plies, competition, predation, life history (Mackay,
1992) and season (Williams, 1980). Williams & Hynes
(1976) showed that four main sources of animals con-
tributed to the recolonization of denuded substrates.
These were vertical movements from within the sub-
strate, aerial sources (e.g. oviposition), upstream and
downstream migration (drift). Downward movement
toward the hyporrheic zone plays an important role
especially in the recolonization of temporary streams
(Delucchi, 1989). The upstream flight of ovigerous
lotic insects is also a principal component of the col-
onization cycle of many taxa (Bird & Hynes, 1980;
Müller, 1982). Drift is the downstream transport of
aquatic organisms in the current. There are different
types of drift, which present seasonal and diel period-
icity patterns and differ in qualitative and quantitative
characteristics (Brittain & Eikeland, 1988). Upstream
movement within the water has been recorded for
many benthic invertebrates (Bishop & Hynes, 1969;
Söderström, 1987). In permanent streams, all four
sources contribute substantially, but positive rheotaxis
and especially drift appear to be the primary com-
ponents. Several studies have described recolonization
mechanisms in North American, northern European
and tropical streams, but until now there have been no
similar investigations in Italian lotic environments.