Do exotic pine plantations favour the spread of invasive
herbivorous mammals in Patagonia?
MARÍA-VICTORIA LANTSCHNER,
1
* VERÓNICA RUSCH
1
AND JOHN P. HAYES
2
1
Grupo de Ecología Forestal, INTA EEA Bariloche, CC 277, 8400 Bariloche, Río Negro, Argentina
(Email: mvlantschner@bariloche.inta.gov.ar); and
2
Department of Wildlife Ecology and Conservation,
University of Florida, Gainesville 32611, Florida, USA
Abstract Changes in land use patterns and vegetation can trigger ecological change in occupancy and commu-
nity composition. Among the potential ecological consequences of land use change is altered susceptibility to
occupancy by invasive species. We investigated the responses of three introduced mammals (red deer, Cervus
elaphus; wild boar, Sus scrofa; and European hare, Lepus europaeus) to replacement of native vegetation by exotic pine
plantations in the Patagonian forest-steppe ecotone using camera-trap surveys (8633 trap-days).We used logistic
regression models to relate species presence with habitat variables at stand and landscape scales. Red deer and wild
boar used pine plantations significantly more frequently than native vegetation. In contrast, occurrence of European
hares did not differ between pine plantations and native vegetation, although hares were recorded more frequently
in firebreaks than in plantations or native vegetation. Presence of red deer and wild boar was positively associated
with cover of pine plantations at the landscape scale, and negatively associated with mid-storey cover and diversity
at the stand scale. European hares preferred sites with low arboreal and mid-storey cover. Our results suggest that
pine plantations promote increased abundances of invasive species whose original distributions are associated with
woodlands (red deer and wild boar), and could act as source or pathways for invasive species to new areas.
Key words: Cervus elaphus, invasion, land use change, Lepus europaeus, Sus scrofa.
INTRODUCTION
Transformation of landscapes by humans may facili-
tate the spread of invasive species, by changing the
spatial and temporal distribution of resources (Sakai
et al. 2001; With 2002). In general the replacement of
native vegetation by exotic conifer plantations, results
in an impoverished flora and fauna (Hartley 2002;
Brockerhoff et al. 2008); however, some species,
including some invasives, can benefit from the habitat
provided in these altered habitats (Allen et al. 1995;
Odgen et al. 1997). Moreover, synergy among non-
indigenous species could facilitate invasions and
increase the likelihood of survival and occurrence and
magnitude of ecological impact, as predicted under the
invasional meltdown hypothesis (Simberloff & Von
Holle 1999).
Exotic forest plantations often have been found to
favour invasion of exotic species, including plants
(Allen et al. 1995; Chiarucci & De Dominicis 1995;
Odgen et al. 1997; Hobbs et al. 2003; Ito et al. 2004;
Paritsis & Aizen 2008), arthropods (Neuman 1979;
Bonham et al. 2002; Pawson et al. 2008), birds (Clout
& Gaze 1984; Lindenmayer et al. 2002), and mammals
(Barnett et al. 1977; Lindenmayer et al. 1999). The
increased abundance of exotic species in exotic forest
plantations may result from increased disturbance,
changes in light and soil conditions, changes in land
management, and increased availability of certain
resources (Bremer & Farley 2010).
In Argentinean Patagonia, exotic fast-growing plan-
tations have been established since the 1970s in the
forest-steppe ecotone, strongly promoted by the state
(Schlichter & Laclau 1998). There are approximately
80 000 ha of planted forests, mainly Pinus ponderosa,
and to a lesser degree, Pseudotsuga menziesii and
Pinus contora (Loguercio & Deccechis 2006; CFI-
FUNDFAEP 2009). A few herbivorous mammal
species also were introduced into Patagonia during the
late 1800s and early 1900s, mainly for hunting
(Grigera & Rapoport 1983; Jaksic et al. 2002). The
most widespread species are European hare (Lepus
europaeus), red deer (Cervus elaphus), and wild boar
(Sus scrofa). While these species benefit local econo-
mies (Novaro & Walker 2005), they may affect native
ecosystems in several ways (Vázquez 2002). Through
browsing, grazing and trampling, these species have
altered floristic composition and stand structure in
Andean rain forests and xeric woodlands (Veblen et al.
1992; Vázquez 2002).These introduced herbivores are
also suggested to affect native herbivores through
*Corresponding author.
Accepted for publication April 2012.
Austral Ecology (2012) ••, ••–••
© 2012 The Authors doi:10.1111/j.1442-9993.2012.02411.x
Austral Ecology © 2012 Ecological Society of Australia