Journal of species lists and distribution
Chec List
487
L ISTS OF S PECIES
Check List 9(3): 487–503, 2013
© 2013 Check List and Authors
ISSN 1809-127X (available at www.checklist.org.br)
In South America, the largest savanna region is located
in Brazil, and is called Cerrado, and as in other savannas,
the fire regime is an important factor in the evolution
of the landscape and, consequently, of the vegetation
(Gottsberger and Silberbauer-Gottsberger 2006a). In
savannic physiognomies the presence of trees is greater
in areas protected from fire, especially in dry-shrub
savannas, where protection allows the regeneration of the
woody component (Moreira 2000). In general, the absence
of fire benefits the woody component and increases the
structural complexity of vegetation, while the passage of
fire benefits the non-woody component and increases the
presence of herbs and subshrubs in the landscape (Mistry
1998).
In the Cerrado, most of the ongoing studies on
dynamics have focused only on the tree layer (Libano
and Felfili 2006; Aquino et al. 2007; Roitman et al. 2008,
Carvalho and Felfili 2011). Studies that directly focus on
understanding the temporal dynamics of the herbaceous
and shrub layer over the years are still scarce, especially
with regard to the grassland physiognomies of the biome
(Eugênio et al. 2011).
The Cerrado phytogeographical domain has a very
heterogeneous physiognomy, that ranges from open
grasslands to dense forests, but which has as the most
common formation the savannic physiognomy, known
as cerrado sensu stricto (Oliveira-Filho and Ratter 2002;
Ab’Sáber 2003; Ribeiro and Walter 2008). The grassland
formations include wet grassland (campo limpo), dry-
shrub grassland (campo sujo) and “campo rupestre”
(rupiculous field grassland) (Ribeiro and Walter 2008),
and until 2007 these occupied 7% of the entire Cerrado. In
the Federal District alone, they covered a total of 6,164 ha
(Sano et al. 2007).
The dry-shrub grassland (Campo sujo) is one of the
Introduction
The dynamic process in a community is characterized
by patterns, mechanisms and, in many systems, successive
disturbances are important sources of changes in the
landscape (Glenn-Lewin and van der Maarel 1992). The
progression of changes in the composition and structure
of a community over time, due to disturbances in the
environment, is conceptualized as a succession process or
directional change (Buchanan 1982).
Studies of temporal dynamics for grassland sites report
that the suppression of disturbances such as fire plays
a crucial role in the floristic and structural changes of
these communities (San José and Fariñas 1991; Moreira
2000; Behling et al. 2007; Kahmen and Poschlod 2008;
Ravi and D’Odorico 2009). Changes over time are related
to differences in species abundance and composition,
differences in the spectrum of life forms and functional
characteristics. The suppression of fire in savannas
intervenes in natural ecological processes, and in more
open areas like grasslands, a gradual increase in the density
of woody and fire-sensitive species can be seen (San José
and Fariñas 1983; 1991; Silva et al. 2001; Durigan and
Ratter 2006; Gardner 2006; Pinheiro and Durigan 2009).
Savannas are considered dynamic ecotones, distributed
between grassland formations and more densely
vegetated areas (Coutinho 1978, Roitman et al. 2008). In
these landscapes grasses and trees coexist, influenced by
interactions with the climate, soil and disturbances such as
fire, and fluctuations in any of these factors may result in
an increase in certain life forms (Roitman et al. 2008). The
intensification or suppression of disturbances modifies the
composition of species in an area. The landscape is altered
by the exclusion of sensitive species in the first case, and
by the exclusion of resistant species in the second (Libano
and Felfili 2006).
Abstract: Studies of temporal dynamics for grassland sites report that fire suppression plays a crucial role in floristic
changes. The objective of this study was to verify whether after seven years without fire, communities showed variations in
terms of composition, life forms, pollination and dispersal syndromes. The first survey (T0) was conducted from September
1999 to October 2000, while the second (T1) took place from August 2006 to August 2007. The floristic results in T1
were compared with the survey in T0 through the Sorensen similarity index and Chi-square tests. Over time, there were
differences in the composition, life forms and pollination and dispersion syndromes. The evidence of changes suggests that
the frequency of the fire regime can be considered the main agent for change in the flora of these communities.
1 Universidade de Brasília, Instituto de Biologia, Departamento de Botânica. Campus Darcy Ribeiro, Instituto de Ciências Biológicas, bloco D, 1°
piso. CEP 70910-900. Asa Norte, Brasília, DF, Brazil.
2 Universidade Católica de Brasília, Curso de Biologia, Laboratório de Botânica. Campus I - QS 07 Lote 01 EPCT, Águas Claras. CEP 71966-700 –
Taguatinga, DF, Brazil.
3 Universidade de Brasília, Departamento de Engenharia Florestal. Campus Darcy Ribeiro, Brasília, DF, Brazil. In memoriam
* Corresponding author. E-mail: aryanne_amaral@yahoo.com.br
Aryanne Gonçalves Amaral
1*
, Cássia Beatriz R. Munhoz
1
, Chesterton Ulysses Orlando Eugênio
1,2
and
Jeanine Maria Felfili
3
Vascular flora in dry-shrub and wet grassland Cerrado
seven years after a fire, Federal District, Brazil