SYSTEMATICS Two New Species of Aphids Found in Lebanese Amber and a Revision of the Family Tajmyraphididae kononova, 1975 (Hemiptera: Sternorrhyncha) OLE E. HEIE 1 AND DANY AZAR 2 Ann. Entomol. Soc. Am. 93(6): 1222Ð1225 (2000) ABSTRACT Two new species of fossil aphids found in Lebanese amber are described, namely Megarostrum azari Heie n. gen., n. sp. and Lebanaphis minor Heie n. gen., n. sp. Both have a very long rostrum and are in this respect different from previously described species of the family Tajmyraphi- didae, which is subdivided into Þve new subfamilies. KEY WORDS Lebanon, amber, fossil, aphids, Lower Cretaceous TWO PIECES OF Lebanese amber with alate aphids were found by D.A. and sent for identiÞcation to O.E.H. Both specimens belong to previously undescribed and extinct species. The amber pieces are from the locality of Mdeyrij/ Hammana in Caza Baabda, central Lebanon. The age is Lower Cretaceous, Late Neocomian/Lower Aptian, 120 Ð125 Myr. They were prepared by D.A. on mi- croscope slides in a Canada balsam medium under cover glasses. They were studied by the senior author using a Zeiss binocular microscope (Gu ¨ ttengen, Germany) with a 10 magniÞcation. The thickness of the material pre- vented the use of a stronger magniÞcation. The two species were both well preserved, and it was possible to see them both from the dorsal and ventral side. The amber was light brown to yellowish and quite clear. The aphids were studied by light coming from beneath and light coming from above. All measurements are given in millimeters. These species belong to the family Tajmyraphididae as diagnosed by Kononova (1975) and represent two new genera. The diagnosis of this family is as follows: pterostigma short; radial sector leaving the distal part of the pterostigma; media departing from its base; cubitus-branches separated at bases, the distal branch leaving the main vein at an angle of 35Ð50°, the prox- imal branch short and forming an angle of 75Ð90° with the main vein; antennae short and rather thick, 4 Ð 6 segmented; siphunculi absent; ovipositor present. It can be added that some species described after 1975 have seven antennal segments, that their processus terminalis is little developed or absent, their pterostigma is rather short and broad, and the tarsi in some species are relatively long. This family has hitherto only been represented by eight species from the Upper Cretaceous, seven from Siberian amber and one from Canadian amber. Megarostrum Heie n. gen. Type Species. Megarostrum azari Heie n. sp. Diagnosis. Rostrum extremely long, more than twice as long as the body. Proximal branch of cubitus of the fore wing very weakly pigmented. Etymology. Mega = very large; species with a very long rostrum. Megarostrum azari Heie n. sp. Material. Holotype No. 476, collector D.A., depos- ited in Muse ´ um National dÕHistoire Naturelle, Paris. Etymology. The speciÞc name is made after D.A., who found the aphid. Stratum typicum. Lower Cretaceous, Late Neoco- mian/Lower Aptian, 120 Ð125 Myr. Locus typicus. Mdeyrij/Hammana in Caza Baabda, central Lebanon. Description. Alate specimen: body 1.2 mm long, with head, most of thorax, sclerites on abdomen and appendages dark pigmented. Hairs not visible except on rostrum, but probably present. Width of head across eyes 0.36 mm; frons straight with protruding ocellus; longitudinal diameter of compound eye 0.10 mm; ocular tubercles apparently present, but difÞcult to distinguish; antenna 0.43 mm long, one-third of the body length, 7-segmented, rather thick; length of an- tennal segments I = 0.03 mm, II = 0.07 mm, III = 0.14 mm, IV = 0.05 mm, V = 0.04 mm, VI = 0.04 mm, VII = 0.07 mm; a processus terminalis not visible. Rostrum longer than twice the body length, apex broken, the present part 3.0 mm long, reaching 2.10 mm behind the posterior end of the body and very slender, the present part of the apical segment 1.37 mm long, 0.04 mm wide, with numerous rather thick and pointed 1 Biology Department, DLH, Emdrupvej 101, DK-2400 Copenha- gen NV, Denmark (e-mail: tron@dlh1.dlh.dk). 2 Muse ´ um National dÕHistoire Naturelle, Laboratoire dÕEntomologie, 45, Rue Buffon, F-75005 Paris, France (e-mail: azar@mnhn.fr). 0013-8746/00/1222Ð1225$02.00/0  2000 Entomological Society of America